Lianas, woody, usually densely rust-colored hairy. Leaves opposite, rarely subopposite, petiolate, entire, papery, with pinnate veins, with tendrils between leaf pairs. Plants dioecious. Cymose panicles axillary or extra-axillary. Flowers small; pedicel jointed. Staminate flowers: calyx cupular, 5-toothed; corolla deeply (3 or)4-or 5-lobed, coherent at base, outside densely hairy; stamens 3-5, alternate with corolla lobes; filaments broad, short, rarely absent; reduced ovary small or absent. Pistillate flowers: calyx as in staminate flowers, persistent; corolla 4-or 5-lobed, base tubular and often inflated; reduced stamens absent; ovary sessile or with short pedicel, 1-loculed; ovules 2, pendulous from apex; stigma thick, peltate, apex concave, sometimes slightly oblique. Drupe obliquely obovoid, with persistent calyx; exocarp thinly crustaceous; mesocarp thin; endocarp outside reticulate polygonous concave. Seeds with fleshy endosperm; cotyledons flat, leafy.
Lianes with tendrils arising at leaf nodes. Leaves opposite, entire, penninerved. Flowers borne in axillary cymose panicles, rarely terminal umbels, dioecious, pedicellate, articulated below calyx. Sepals partially united to give 4–5 deltoid lobes. Petals 3–5, ± united at the base. Male flowers with 3–5 stamens alternating with petals; filaments very short, usually flattened. Female flowers: ovary subsessile, unilocular; stigma sessile, discoid; ovules 2, pendulous. Fruit a drupe, ellipsoid; endocarp woody; seed large.