Rhizome dorsiventrally flattened, 1.5-3.5 mm wide, long-creeping, internodes to 10 cm long, not white waxy, roots absent, hairs in a ventral band on the rhizome; vascular strands 7-15, with sclerified sheaths, sclerenchyma strands absent. Rhizome scales peltate, sparsely set, distinctly spreading, narrowly ovate or triangular, 0.5-2 by 0.1-0.2 mm, clathrate or subclathrate, margin denticulate. Fronds strongly dimorphic, stipitate, thin-herbaceous. Fertile fronds: lamina simple, linear, 15-30 by 0.1-0.5 cm; stipe 2-7 cm long. Sterile fronds: stipe 2-9 cm long, 0.9-1.5 mm thick, narrowly winged for a considerable part, lamina simple, narrowly ovate to elliptic, 9-36 by 1.1-6.5 cm, index 3-10, base cuneate to narrowly angustate, sometimes cordate and auriculate, margin entire, lower surface without acicular hairs. Venation: veins prominent and distinct, 7-10 mm apart, zigzag, branching at or near the costa, each costal areole giving rise to two veins, connecting veins 2-4, not clearly differentiated, cata-or anadromous, mostly forming several rows of elongated ± hexagonal areoles, which become gradually smaller towards the margin, the costal areole bordered by several areoles, included veins prominent and distinct, free veinlets simple or once forked. Sori acrostichoid, covering the lamina.
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Rhizome 1.5-3.5 mm in diam., dorsiventrally flattened, bearing scales and hairs, with only circumvascular sheaths; vascular bundles 7-15; roots absent, root hairs on rhizome; scales sparse, peltate, distinctly spreading, narrowly ovate or triangular, broadest at middle, 0.5-2 × 0.1-0.2 mm, margin denticulate; phyllopodia 3-80 mm apart, ± distinct. Sterile fronds: stipe 2-9 cm, 0.9-1.5 mm in diam.; lamina narrowly elliptic or narrowly ovate, 9-36 × 1.1-6.5 cm, 3-10 × as long as broad, with short glandular hairs, base narrowly decrescent, stipe winged for a considerable part to cuneate-decrescent to cuneate to cordate, auriculate, margin entire. Fertile fronds: stipe 2-7 cm; lamina simple, linear, 15-30 × 0.1-0.5 cm; veins 7-10 mm apart, prominent and distinct, zigzag, each costal areole giving rise to 2 lateral veins, thus lateral veins seemingly branching at or near costa, costal areole bordered by several smaller areoles; a prominent basiscopic (or sometimes acroscopic) connecting vein dichotomously branching off near costa; connecting veins catadromous, 2-4 between adjacent secondary veins; smaller veins prominent and distinct; free veinlets simple or once forked, usually in-and excurrent. Sori acrostichoid, superficial or slightly sunken; paraphyses present.
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