Rhizome creeping, thick, 3-10 mm in diam., dorsiventrally flattened or subcylindrical, not white waxy. Scales pseudopeltate, ovate or triangular, 1.5-9 × 1-3 mm, margin entire, apex acute, clathrate except for hyaline marginal region, central region bearing multiseptate hairs at least when young. Fronds not or slightly dimorphic. Stipe up to 15 cm, 3-5 mm in diam. Lamina simple, ovate to elliptic or narrowly so to linear, (5-) 25-110 × (1-)5-15 cm, membranous, base narrowly decrescent, stipe winged for a considerable part, margin entire, apex acuminate. Veins prominent and distinct. Sori separate, on whole surface of lamina, sometimes forming 2-8 irregular rows between veins or some connate, elongate on veinlets, orbicular or elongate, 1-2 mm in diam., or length ca. 2.5 mm, superficial or slightly sunken.