Rhizome terete, 1.5-5 mm thick, long-creeping, internodes 15-60 mm long, not white waxy, roots densely set; vascular strands 7-11, with sclerified sheaths, sclerenchyma strands 50-100. Rhizome scales pseudopeltate, densely set, slightly to distinctly spreading, narrowly ovate or triangular, 3-7 by 1-1.5 mm, margin entire to denticulate, apex acute, clathrate or subclathrate, central region glabrous or bearing multiseptate hairs at least when young. Fronds not or slightly dimorphic, stipitate, herbaceous. Stipe 0.5-15(-25) cm long, 1.5-3(-3.5) mm thick; lamina simple or irregularly lobed, narrowly elliptic to linear, 25-70(-90) by 3-8 cm, index (5.5-)7.5-11(-15.5), base narrowly angustate, decurrent to a long wing, margin entire, sinuate or irregularly lobed, apex acuminate, lower surface without acicular hairs. Venation: veins prominent and distinct, 7-11 mm apart, more or less straight, dichotomously branched near the margin; connecting veins 4 or 5, mostly catadromous, forming equally sized, sometimes irregularly shaped, areoles, sometimes also a row of small primary costal areoles; included veins more or less immersed and indistinct, variously anastomosing, free veinlets simple or once forked. Sori separate, round or elongate along the veinlets, superficial or slightly immersed, 1-2 by 1.5-3 mm, mostly irregularly scattered on the smallest veinlets, 1 or 2 per veinlet, 15-30 per sq.cm, on the whole surface of the lamina or absent from the basal parts for half the total length of the lamina, present or absent in marginal areoles, generally present in costal areoles; paraphyses simple uniseriate hairs with glandular topcells, 2-or 3-celled.