Rhizome dorsiventrally flattened, 1-4 mm wide, short-creeping, internodes up to 11 mm long, not white waxy, roots sparsely set; vascular strands 5-10, with conspicuous sclerified sheaths, sclerenchyma strands absent. Rhizome scales pseudopeltate, densely set, slightly spreading, ovate, narrowly ovate or triangular, (0.5-)1-3 by 0.5-1 mm, margin denticulate, apex acute, clathrate or subclathrate, central region glabrous. Fronds not or slightly dimorphic, stipitate, herbaceous to subcoriaceous. Stipe 0.5-4.5 cm long, 0.5-1.5 mm thick; lamina simple, linear, 25-45 by 0.5-1 cm, index 30-65, base truncate, margin entire, apex acuminate, lower surface with sparse to dense acicular hairs, especially around the sori. Venation: veins immersed and distinct, 5-13 mm apart, more or less straight, dichotomously branched below the middle to near the margin, a single connecting vein forming one row of large areoles parallel to the costa, bordered by several smaller areoles; costal areole, if present, formed by smaller veins; included venation more or less immersed and indistinct, variously anastomosing; free veinlets simple to twice forked. Sori separate, round or elongate, superficial or slightly immersed, 0.5-1.5 by 0.5-40 mm, mostly irregularly scattered on the smallest veinlets, 10-60 per sq.cm, on the whole surface of the lamina, present in marginal areoles, present or absent from costal areoles, sometimes concentrated near the margin or in a marginal longitudinal coenosorus; paraphyses simple uniseriate hairs with glandular topcells, 2-or 3-celled.