Shrubs, lianas or trees. Leaves opposite, often lobed. Capitula many, in terminal panicles; involucre 1–2-seriate, the outer phyllaries leafy, the inner scarious; paleae scarious. Ray florets neuter, white; inner florets hermaphrodite, with auriculate anthers and recurved style branches. Achenes usually compressed, black, glabrous; pappus absent.