Shrubs or trees, to 12 m. Leaves alternate (distichous), petiolate, stipitate; stipules subulate or filiform [absent or peltate discs]; blade palmately veined, seldom lobed, base often asymmetric, margins serrate, surfaces hairy, hairs usually mixed: unicellular and multicellular, simple with some setiform and some glandular, branched, and stellate, often ± tangled, forming tomentum. Inflorescences: flowers solitary or in clusters of 2–3+, usually supra-axillary; involucel absent [bracteoles ca. 15, filiform]. Flowers: sepals caducous [persistent], (4–)5(–7), valvate, basally distinct or weakly connate; petals caducous, (4–)5(–7), distinct; nectaries absent; stamens 10–75+, filaments distinct or bases connate; ovary superior [inferior], 5–7-carpellate; style 1 [0]; stigmas 5–7, ± decurrent. Fruits baccate, ± spheric. Seeds [25–]100–200+.