Orthilia Raf.

Herbs or subshrubs, chlorophyllous, autotrophic. Stems decumbent or erect, glabrous or papillose, especially distally. Leaves cauline, alternate or pseudoverticillate in (1-)2-4 whorls; petiole present; blade not maculate, ovate to broadly ovate or elliptic to orbiculate, subcoriaceous, margins entire or undulate to serrulate, rarely serrate, plane or, rarely, obscurely revolute, surfaces glabrous. Inflorescences racemes, often lax in bud or flower, usually erect in fruit, (secund); peduncular bracts usually present; inflorescence bracts free from pedicels. Pedicels recurved to reflexed in fruit; bracteoles absent. Flowers radially symmetric, spreading or nodding; sepals 5, connate proximally, often obscurely so, calyx lobes broadly triangular to ovate; petals 5, distinct, greenish white, yellowish white, or white, with 2 inconspicuous basal tubercles, corolla suburceolate; intrastaminal nectary disc absent; stamens 10, included or barely exserted; filaments broad proximally, gradually narrowed medially, slender distally, glabrous; anthers oblong, without awns, without tubules, dehiscent by 2 round to elliptic pores; pistil 5-carpellate; ovary imperfectly 5-locular; placentation intruded-parietal; style (exserted), straight or nearly so, expanded distally or not; stigma 5-ridged, without subtending ring of hairs. Fruits capsular, pendulous, dehiscence loculicidal, cobwebby tissue exposed by splitting valves at dehiscence. Seeds ca. 1000, fusiform, winged. x = 19.