Trees or sometimes shrubs, monoecious. Indumentum of solitary simple hairs only. Leaves imparipinnate or (pari)pinnate, 1-6-jugate; no pseudo-stipules; petiole and rachis terete, not winged. Leaflets smooth beneath; the margin entire to dentate; nervation open, when the margin is dentate each second nerve is ending in a tooth. Inflorescences ramiflorous, axillary, or terminal. Flowers unisexual, regular. Sepals (4 or) 5(-7), slightly connate to free, all equal (to very unequal), not petaloid, outside white to dark yellow hairy, margin ciliate, inside white sericeous, no glands, margin entire. Petals (4 or) 5(-7), longer than the sepals, distinctly clawed to broadly truncate at base, outside usually glabrous to laxly woolly, inside glabrous; blade variably developed to nearly completely reduced; scale usually larger than the blade, emarginate to divided into two lobes, both sides orange woolly especially at the upper margin; crest absent. Disc composed of a flat ring adnate to the torus except for the margin, thin fleshy, glabrous, margin with an erect rim to tubular collar up to 1 mm high. Stamens 5-8 (or 9), distinctly exserted in male flowers, glabrous; dehiscence latrorse. Pistil sessile; ovary densely tuberculate, each tubercle bearing a stiff erect hair, further glabrous to minutely hairy, (l-)3-(or 4-)locular; ovules 1 per locule; style apical, longer than the ovary, laxly to densely short-strigose; stigma flat to with recurved lobes up to 1.5 mm long. Fruits capsular, ± globular, sessile, not winged, smooth via rough, ribbed, or warty to densely spiny, loculicidally dehiscing with 3 or 4 mostly unequal valves or tearing apart at random; wall thick, fibrous-woody; hairy inside. Seeds usually 1 per fruit only, about globular to slightly 2-or 3-lobed; no arillode, but the white hilar spot covering up to ¾ of the seed; the membranous remains of the septae with the non-developed ovules are tightly pressed against the seed.
More
Shrubs or trees, monoecious or dioecious. Leaves imparipinnate or (pari)pinnate, petiolate; leaflets 1-5-jugate, leathery, margin entire or sparsely dentate. Thyrses axillary or terminal; bracts and bracteoles small. Flowers actinomorphic, unisexual. Calyx cupular, 5-lobed; sepals triangular, valvate. Petals 5, small, slightly longer than calyx, slender, with 1 large scale adaxially; scale connate with margin of petals funnelform. Disk annular, 5-lobed. Stamens (male flowers) 6-10, exserted; filaments glabrous or hairy. Ovary (female flowers) obovoid, tuberculous; ovules 1 per locule; style terminal; stigma 3-lobed, lobes excurved. Capsules subglobose, usually rudimentary 1-loculed, loculicidal into 3 schizocarps; pericarp leathery or fibrous-woody, tuberculous or woody-spiny. Seeds compressed-globose or transversely ellipsoid; testa leathery; hilum transversely elliptic, large; embryo arched.
Home
