Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4-6(-10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). Stems simple from base, erect, (usually unbranched), to 4.5 dm. Basal leaves: blade suborbicular to elliptic, 2-9.5(-11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. Cauline leaves similar to basal, blade narrowly linear or, sometimes, orbicular. Racemes loose (lax). Fruiting pedicels (recurved, straight, curved, or sigmoid), 6-20 mm. Flowers: sepals elliptic or ovate, (2-)3.5-7.5 mm, (cucullate); petals 6-11 mm. Fruits obovoid to subglobose, not inflated (not angustiseptate), 3-6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4 (-8) per ovary; style (1.6-)3-6 mm. Seeds flattened. 2n = 10, 30.
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