Annuals, biennials, or perennials; (caudex often present, enlarged, usually branched); not scapose; usually pubescent, trichomes usually sessile, sometimes subsessile or shortly stalked, usually stellate, sometimes stellate-scalelike, rarely simple. Stems erect, spreading, decumbent, or prostrate, unbranched or branched distally. Leaves basal and cauline; petiolate or sessile; basal usually rosulate, petiolate, blade margins usually entire, sometimes repand to pinnatifid; cauline petiolate or sessile, blade margins usually entire, sometimes repand to dentate. Racemes (few-to several-flowered, proximalmost flowers rarely bracteate), elongated or not in fruit. Fruiting pedicels erect, horizontal, divaricate, spreading, ascending, or sigmoid, usually slender, sometimes stout. Flowers: sepals erect or spreading, linear, lanceolate, elliptic, oblong, ovate, or deltate, lateral pair usually saccate basally, sometimes subsaccate or not saccate; petals usually yellow, sometimes orange (occasionally drying purplish or maroon), rarely white or purple, spatulate, obovate, ovate, oblanceolate, or obdeltate, (longer than sepals), claw differentiated or not from blade, (apex usually rounded, rarely slightly emarginate); stamens tetradynamous; filaments usually not dilated basally; anthers ovate to narrowly oblong, (apex usually obtuse); nectar glands confluent, subtending bases of stamens, median glands present or absent. Fruits silicles, sessile, subsessile or, rarely, shortly stipitate, globose, subglobose, orbicular, suborbicular, ellipsoid, elliptic, lanceolate, obcordate, obdeltate, oblong, obpyriform, ovate, ovoid, or obovoid, not torulose, inflated or not, terete, latiseptate, or angustiseptate; valves each often with obscure midvein, (usually not retaining seeds after dehiscence), usually pubescent, sometimes glabrous, or, rarely, pubescent inside; replum rounded to narrowly oblong; septum usually complete, sometimes perforate, or, rarely, reduced to a rim (often with apical midvein extending to center); ovules (2-)4-32(-40[-80]) per ovary; style distinct; stigma entire. Seeds biseriate, often flattened, sometimes plump, rarely lenticular, usually not winged, rarely narrowly winged or margined, often suborbicular; seed coat (smooth), mucilaginous or not when wetted; cotyledons accumbent. x = 4, 5, 6, 7, 8, 9, 10, 12, 15.
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