Perennials; caudex branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays mostly furcate, (arms of unequal lengths, finely tuberculate). Stems several from base, erect or lateral decumbent, (unbranched), 1-2 dm. Basal leaves: (petiole usually winged); blade broadly oblanceolate, 3-8 cm, margins usually dentate or pinnatifid, rarely subentire, (terminal lobe acute or obtuse, not rounded). Cauline leaves: blade spatulate to linear-oblanceolate, 1-3 cm, margins usually entire, rarely toothed, (apex acute). Racemes loose (and greatly elongated in fruit) to congested. Fruiting pedicels (recurved), 6-15 mm. Flowers: sepals linear-oblong, 5-7 mm; petals spatulate, 9-11 mm. Fruits (usually pendent on arching pedicels, less frequently widely divergent), irregular in shape, (base obtuse or slightly cordate, apex deeply and broadly notched), not strongly inflated, 8-11 × 8-12 mm, (papery; valves retaining seeds after dehiscence; replum linear-oblong, constricted, 2.5-4 mm, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style 5-8 mm. Seeds flattened.