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Caudex short, erect or suberect; scales firm, narrow, entire with cordate base, multicellular hairs present with the scales and variously on the fronds; fronds simple-pinnate; external form of rachis and its relationship to pinnae as in Davallia; pinnae deeply lobed, the lobes ± crenate or dentate; veins all free, once or twice forked, the basal basiscopic vein in each lobe springing from the base of the costule (rarely from the costa), the basiscopic branch of the basal acroscopic vein (or sometimes both branches) passing to a tooth in the sinus between that lobe and the next, the tooth projecting out of the plane of the lamina; unicellular glands absent on the lamina and from sporangium-stalks; sori on the acroscopic branch of each vein terminal or not, indusiate, indusia reniform, sometimes asymmetric.
Terrestrial or lithophytic ferns; rhizome erect, covered by persistent stipe bases. Stipe conspicuously grooved; vascular strands several, small. Fronds suberect, mostly bipinnatisect; pinnae and the apical pinna usually all similar, occasionally the basal pinnae with elongated pinnules (not on Christmas Island); sinuses between pinna lobes with a small, prominent tooth at base; veins all free. Sori circular, small, on lamina beneath, usually at apex of veinlets in lobes; indusium reniform; paraphyses absent.
Vascular bundles in stipes in a simple ring; veins free
