Pycnarrhena Miers ex Hook.F. & Thomson

Lianas or scandent shrubs. Leaves with petioles swollen at the base and usually at apex, leaving a prominent cup-like scar on the stem; lamina usually penninerved and ± elliptic, sometimes (in Mal., P. insignis) with lower nerves crowd-ed at base and ovate. Inflorescences axillary or ramiflorous, mostly cymose, the peduncles sometimes 1-flowered and fascicled. Male flowers: sepals 6-15 in whorls of 3, imbricate, the outer ones minute, inner ones largest (in P. ozantha the innermost sometimes smaller), rotund and concave; petals 3-6, or 0 in P. ozantha, minute, mostly broadly obovate; stamens variable in number, 6-18, tightly clustered with the filaments usually connate for most of their length (shortly connate in P. lucidd). Female flowers: sepals and petals as in male; staminodes 0; carpels 3-6 with recurved stigmas. Drupes ± subglobose with style-scar on the ventral side below the apex, arising from the margin of a swollen ± globose carpophore; endocarp usually thin, papyraceous to crustaceous and smooth to wrinkled, but in P. ozantha bony and verrucose; endosperm absent; seed broadly ellipsoidal, cotyledons large and thick, very slightly curved.