Sanguisorba L.

Burnet (en), Pimprenelle (fr), Sanguisorbe (fr)

Genus

Angiosperms > Rosales > Rosaceae

Characteristics

Herbs, perennial, 2–20 dm; rhizomatous. Stems 1–10+, ascending to erect, glabrous or glabrescent. Leaves deciduous, basal and cauline; stipules persistent, adnate to petiole, free portion usually ovate, obovate, or suborbiculate, margins dentate; petiole present; blade oblong-elliptic, 5–55 cm (basal blades largest, cauline reduced distally), herbaceous, leaflets (3–)7–21, orbiculate, ovate, or oblong, ?leafy stipels sometimes present on vigorous leaves?, margins flat, crenate or serrate, surfaces glabrous or sparsely hairy. Inflorescences terminal or axillary to distal leaves, 50–500+-flowered, spikes, ?ellipsoid to cylindric?, glabrous or variously pubescent; peduncles present; bracts absent; bracteoles present. Pedicels absent. Flowers 2–5 mm diam.; hypanthium urceolate, 1–6 mm, ?smooth, muricate, or winged?, glabrous; sepals 4, ?basally connate?, spreading, petaloid, elliptic to ovate; petals 0; stamens [2–]4[–12], shorter or longer than sepals; carpels 1[or 2], glabrous, ?styles repeatedly branched, brushlike; ovules pendulous?. Fruits achenes, 1, globose, 0.5–2 mm, glabrous; hypanthium persistent, enclosing achenes, ?dry, hardened, top-shaped to ellipsoid, 2–5 mm, 4-angled or-winged, faces smooth between angles?; sepals persistent, spreading to ascending. x = 14.
More
Herbs perennial. Rootstock robust, bearing many fusiform, cylindric roots in lower part. Stipules sheathing, adnate to petiole, petiole sheathing and imbricate at base; leaf blade imparipinnate; leaflets serrate at margin. Inflorescences terminal on elongate scapes, densely capitate or spicate, bracteate and bracteolate. Flowers bisexual, rarely unisexual (when plants monoecious). Hypanthium with a constricted throat. Sepals 4(–7), imbricate, petaloid, purple, red, pink, or white, rarely greenish. Petals absent. Disk lining hypanthium. Stamens usually 4, rarely more, inserted in throat of hypanthium; filaments free, rarely partly connate; anthers didymous. Carpel 1(or 2), included in hypanthium; ovule pendulous; style terminal, filiform; stigma penicillate. Achene dry, included in hardened, muricate or winged hypanthium. Cotyledons planoconvex. x = 7.
Perennial herbs, often arising from thick, branching rhizomes. Lvs alternate, mostly in basal rosettes, imparipinnate; leaflets serrate; stipules persistent, adnate to petiole for lower ⅔. Fls in dense terminal clusters or spikes, subtended by several bracteoles, ☿ or unisexual, 4-merous, small, sessile. Hypanthium deeply concave. Epicalyx 0. Sepals greenish or purplish. Petals 0. Stamens 4-numerous. Ovary superior, surrounded by the hypanthium; carpels 1-2-(3); styles terminal and filiform; ovules solitary. Fr. of 1-(several) achenes enclosed in hardened, dry, 4-angled, sometimes ± flattened or cup-like hypanthium with variously sculptured faces and sometimes thickened or winged angles.
Fls 4-merous, perfect or unisexual; hypanthium urceolate, contracted at the mouth, ± 4-angled, not prickly; sep petaloid; pet none; stamens (2)4 or many; pistils 1 or 2; ovaries included in the hypanthium; ovule 1, suspended; style terminal; fr an achene, enclosed by the indurate hypanthium; perennial herbs from a thick rhizome, or annuals, the lvs pinnately compound with serrate to pinnatifid lfls, the fls small, borne in dense spikes or heads. 25, N. Amer., Eurasia.
Perennial herbs or shrubs, unarmed. Leaves often in rosettes, imparipinnate. Flowers in terminal spikes or heads on usually long and slender peduncles, 4-merous, bi-or unisexual. Hypanthium urceolate, constricted at apex, persistent. Sepals deciduous. Petals absent. Stamens 2-50. Pistils 1 or 2, rarely 3, free; ovaries 1-locular; style terminal, with peni-cillate stigma. Achenes included in hardened hypanthium.
Life form perennial
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality
Pollination -
Spread -
Mature width (meter) 0.5 - 1.0
Mature height (meter) 0.6 - 1.75
Root system rhizome tap-root
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light 4-9
Soil humidity -
Soil texture -
Soil acidity 2-8
Soil nutriment -
Hardiness (USDA) 6-9

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by divisions.
Mode divisions
Germination duration (days) 30 - 60
Germination temperacture (C°) 10 - 11
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -