Securidaca L.

Securidaca (en)

Genus

Angiosperms > Fabales > Polygalaceae

Characteristics

Big lianas to straggling shrubs. Twigs terete, sometimes twisted, glabrescent, at the nodes often with a pair of slightly protruding glands. Leaves alternate or distichous, petioled. Inflorescences raceme-like or usually paniculate, (supra-) axillary or terminal, at the nodes usually with a pair of protruding glands; bracts and bracteoles early caducous. Sepals 5, unequal, caducous before fruit-setting, ciliate, the lateral ones (alae) at least twice as large as the other ones and petaloid, unguiculate, auriculate. Petals 3, halfway adnate to the staminal tube or with an additional pair of reduced petals; upper petals spathulate, lower one (keel) boat-shaped, unguiculate, at apex with or rarely without a slightly 2-lobed, obliquely cup-shaped, hardly incised appendage (crest). Stamens 8, monadelphous; anthers bisporangiate by abortion of the outer microsporan-giae, sessile or on a free filamentous stalk, opening by an apical, oblique pore common to both cells. Disk annular, distinct after flowering. Ovary asymmetrical, 1-celled (by abortion of the second cell), 1-ovuled; style subterminal, curved in apical half, with a terminal, ± 2-lobed stigma. Fruit a samara with a coriaceous wing (wing sometimes reduced), sometimes a second reduced wing present. Seed inappendiculate, glabrous, with thick cotyledons.
More
Woody vines, usually scandent shrubs, or trees. Leaves alternate, entire, short-petioled, generally with corneous peziziform stipular glands. Racemes terminal or axillary, often paniculate. Flowers usually pink or purple; sepals 5, free, the outer 3 herbaceous, the inner 2 (wings) much larger, petal-like; petals 3, the lowest (keel) boat-shaped, clawed, with a plicate, fimbriate crest, the 2 upper adnate to the base of the staminal tube but distinct from the keel; stamens 8, their filaments connate nearly to the apex into a sheath split on the upper side, adnate to the keel and upper petals toward the base; anthers confluently 1-celled, open-ing by a large introrse-apical pore; ovary unilocular through the abortion of the second locule, with an appendage on one side which develops into the wing of the fruit, the style sickle-shaped, excavated apically, the stigma 2-lobed, approxi-mate, the ovule solitary, pendulous; disc a low fleshy ring at base of the ovary. Fruit a samara, the wing large, 1-seeded; seed glabrous, exarillate, endosperm wanting, the cotyledons thick-fleshy, oily.
Shrubs scandent. Leaves simple, alternate; nodal glands present, veinlets reticulate. Inflorescences racemose or paniculate, terminal or axillary. Flowers zygomorphic, small, with bracts. Sepals 5, caducous, unequal; outer 3 small; inner 2 petaloid, large. Petals 3, lateral petals and keel connate or free, keel helmet-shaped, with apical appendage. Stamens 8; filaments united in lower part, forming trough-shaped sheath open on upper side and adnate to petals; anthers 2-celled, ovoid, dehiscent by oblique pores. Disk reniform. Ovary 1-loculed; ovule 1, anatropous, pendulous; style falcate, curved; stigma short, lobed or not. Fruit usually a samara, with 1 seed; wing oblong or rhombic-oblong, leathery, veins numerous. Seed subglobose, without endosperm, without strophiole; seed coat membranous.
Fruit a samara (a 1-locular, 1-seeded, winged nut) usually 1-winged, occasionally with an additional rudimentary wing.
Small trees or shrubs, often scandent, or lianes with entire, alternate leaves.
Sepals 5, unequal, two lateral ones (wing-sepals) larger and petaloid, free.
Petals 3 (occasionally 5, two being vestigial), lowest keel-shaped.
Flowers zygomorphic, in terminal and axillary racemes or panicles.
Seed without caruncle and without endosperm.
Stamens 8, monadelphous.
Ovary 1-locular.
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Growth form shrub
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Foliage retention deciduous
Sexuality hermaphrodite
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Environment

Primary and secondary forests, along streams and in ravines, 0-1500 m.
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Hardiness (USDA) 8-12

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