Polygalaceae Hoffmanns. & Link

Milkwort family (en), Polygalacées (fr)

Family

Angiosperms > Fabales

Characteristics

Perennial or annual herbs or shrubs or trees, rarely small herbs (the latter sometimes saprophytic). Leaves simple, alternate, opposite, or whorled, petiolate or sessile, papery or leathery, with pinnate veins, margin entire, leaves rarely reduced and scalelike; stipules absent, sometimes spiniform or scalelike appendages present. Flowers bisexual, zygomorphic, white, yellow, or purple-red, pedicellate or sessile, in axillary or terminal racemes, panicles, or spikes, with bracts and usually also with bracteoles. Calyx persistent or caducous; sepals 5, free or connate at base, outer 3 small, inner 2 (alae) large, petal-like, or all 5 nearly equal. Petals 3 or 5, basally often connate, lower (median) one ("keel") usually inflexed, carinate, sometimes with fimbriate or lamellate or papilionaceous apical appendages. Stamens 8, 7, 5, or 4; filaments free, or variously united and forming a sheath open on upper side and troughlike; anthers basifixed, usually dehiscing by a single apical pore. Disk usually absent, if present annular or glandular. Ovary superior, 1-or 2-loculed; ovule 1 per locule, anatropous, pendulous, rarely ovules numerous and placentas parietal; style 1, erect or curved; stigmas 1 or 2, capitate. Fruit a 2-loculed capsule, dehiscing by valves, or a 1-loculed samara or a berrylike drupe, dehiscing or not. Seeds 2, or 1 with 1 sterile locule, yellow-brown, dark castaneous, or black, ovoid, globose, or ellipsoidal, glabrous or piliferous, strophiolate or not, with or without endosperm, sometimes with an appendage at end opposite to strophiole.
More
Annual or perennial herbs, climbers, shrubs or trees, rarely saprotrophic (not in Australia); roots often aromatic; indumentum of unicellular hairs or plants glabrous. Leaves alternate, rarely opposite or whorled, simple, sessile or petiolate, sometimes spinose or with nectary glands; stipules absent. Inflorescence usually terminal or axillary, in racemes, panicles or spikes, rarely flowers solitary. Flowers bisexual, zygomorphic, white, yellow, pink, purple or blue, usually subtended by a bract and 2 bracteoles. Sepals 5, free or the lower 2 united or rarely fused in a tube, subequal or the lateral pair enlarged and petaloid. Petals 3 or 5, free or variously united, sometimes fused to the base of the staminal tube, the lowermost petal often keeled, lobed or pouched and of contrasting colour. Stamens (2–) 5–8 (–10), usually in 2 series; filaments usually united at base; anthers dehiscing by apical pores or longitudinal slits. Gynoecium of 2–8 fused carpels. Ovary superior, 2–8-locular, ovules 1 per locule, or unilocular (rarely incompletely bilocular) with 4–40 ovules per locule; style simple, apically bilobed or dilated. Fruit a loculicidal capsule or a nut, samara, drupe or berry, often winged. Seeds glabrous or hairy, sometimes arillate.       
Stamens 8, rarely 5 or 4, monadelphous beyond the middle or rarely free, the sheath split above, often adnate to the petals; anthers erect, 1–2-celled, opening by an apical pore, rarely 2-valved or opening nearly to the base
Ovary free, 2-celled or rarely 1-celled or 3–5-celled; style simple; ovules solitary in each cell, rarely more, pendulous
Petals 3–5, hypogynous, declinate, outer 2 free or united with the lowermost, upper 2 free or minute and scale-like or 0
Petals 3–5; two upper ones free or joined to the lower one (keel); two lateral ones free, often absent or vestigial
Sepals 5; two lateral (interior) ones often petaloid (wings); two anterior ones sometimes joined
Seeds often pilose, with a conspicuous strophiole; endosperm mostly present; embryo straight
Ovary superior, usually 2-locular (1–5), with 1 pendulous ovule in each loculus
Sepals 5, free, imbricate, the two inner larger, often petaloid, wing-like
Stamens usually 5–8 (rarely 4); filaments usually united in a slit tube
Torus small or rarely expanded into an annular disk within the stamens
Seeds often sericeous, usually carunculate; endosperm usually present
Small trees, woody climbers, shrublets or perennial or annual herbs
Flowers hermaphrodite, zygomorphic; pedicels often articulated
Leaves alternate, rarely opposite, simple; stipules none
Leaves usually alternate, simple, entire, exstipulate
Herbs, shrubs, climbers or rarely small trees
Fruit a capsule, samara or drupe
Fruit a capsule, samara or drupe
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Cultivation

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