Xanthophyllum Roxb.

Genus

Angiosperms > Fabales > Polygalaceae

Characteristics

Shrubs or trees. Twigs terete, sometimes with annular or cap-like nodal glands. Axillary buds 2-8, serial, each with 2 bud scales. Leaves alternate (or spiral), sometimes shifted-decussate, stalked, nearly always with glands beneath, mostly near base of blade. Inflorescence axillary, branched or sometimes unbranched, panicle-or raceme-like. Flowers solitary or in lower half with 3 (rarely more) together. Sepals 5, free, usually slightly unequal, usually caducous after flowering. Petals 5, usually unequal, free; lower petal (carina) usually boat-shaped, clawed, apically inappendiculate. Stamens nearly always 8, rarely 7, 9, or 8-10; 4 epipetalous, 2 placed at the base of the carina and ad-nate with it or not, 2 alternipetalous and opposite the lateral sepals; filaments free or connate basally, rarely halfway connate, very rarely triadelphous; anthers tetrasporangiate, opening introrsely with slits. Disk annular. Gynoecium usually as long as androecium; ovary usually shortly stipitate, syncarpous, composed of 2 median carpels, 1-locular or sometimes semi-2-locular; style terminal; stigma slightly bilobed or sometimes peltate; ovules 2-seriate, 4 or (6-)8-c. 20, very rarely c. 40. Fruit indehiscent (or very rarely irregularly 2-valved, not in Mal.), globular or rarely ellipsoid, c. 1-15 cm, usually with a hard pericarp. Seed(s) 1 or 4-20, inappendiculate, glabrous; embryo large, flat or thick; albumen copious to nearly absent.
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Trees or shrubs. Leaves simple, alternate, without stipules, petiolate; leaf blade usually yellow-green when dry, leathery, veinlets reticulate, margin entire. Flowers bisexual, zygomorphic, shortly pedicellate, in racemes or panicles, axillary or terminal; bracts and bracteoles present. Sepals 5, imbricate, inner 2 slightly larger. Petals 5, differing slightly in size, imbricate, sometimes unguiculate, free, keel without apical appendages. Stamens 8; filaments free or 2-4 adherent at base of ovary, other 4-6 adherent at base of petals, ± slightly inflated, pubescent; anthers introrse, usually piliferous at base. Disk annular, fleshy, shorter than ovary. Ovary superior; carpels 2, connate, stalked, 1-loculed, glabrous or piliferous, placentation parietal; ovules 2 to numerous, anatropous; style 1, hairy; stigma capitate, shallowly lobed. Fruit a drupe, globose, fibrous-fleshy when dry, 1-loculed. Seed usually 1; seed coat membranous; caruncle absent; endosperm absent.
Trees or (not in Australia) shrubs. Leaves alternate, simple, usually coriaceous, petiolate. Stipules absent or represented by nodal glands. Inflorescences axillary or terminal racemes, sometimes paniculate, bracteate and bracteolate. Flowers bisexual, usually zygomorphic, hypogynous. Calyx of 5 free sepals, imbricate (quincuncial), the inner usually longer. Corolla of 5 free petals, imbricate, often clawed, unequal or ± similar, the lowest ± folded forming a keel. Stamens 8; filaments free or (not in Australia) basally connate; anthers basifixed, 2-locular, dehiscing by longitudinal slits. Disc intrastaminal. Ovary superior, stipitate, syncarpous with 2 carpels, 1-locular; placentation parietal; ovules 2–many, anatropous; style terminal; stigma capitate or 2-lobed. Fruit indehiscent or irregularly splitting, dry to subfleshy. Seeds 1–many; embryo large; endosperm absent.
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Environment

By far the majority of the species grow in the tropical lowland rain-forest, mostly below 500 m. About 10 spp. (of which 3 extra-Malesian ones) are usually found between 500-1200(-1500) m, but sometimes also lower. About a dozen species are sometimes found in freshwater swamp forests, but they are not confined to that habitat, except 63. X. ramiflorum. Also nearly all of the continental Southeast Asian species seem to be confined to the rain-forest; 60. X. lanceatum is restricted to stream banks and swamps; 62. X. virens occurs in monsoon forests, tolerating rather long dry periods. Otherwise the ecology is for all species 'rain-forest', not specifically repeated.
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Hardiness (USDA) 8-12

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Images

Xanthophyllum unspecified picture

Distribution

Xanthophyllum world distribution map, present in Australia and China

Identifiers

LSID urn:lsid:ipni.org:names:332067-2
WFO ID wfo-4000040889
COL ID 649JB
BDTFX ID -
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Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Skaphium Xanthophyllum

Lower taxons

Xanthophyllum retinerve Xanthophyllum tenue Xanthophyllum tardicrescens Xanthophyllum suberosum Xanthophyllum schixocarpon Xanthophyllum rufum Xanthophyllum rheophilum Xanthophyllum resupinatum Xanthophyllum reflexum Xanthophyllum stipitatum Xanthophyllum tenuipetalum Xanthophyllum zeylanicum Xanthophyllum virens Xanthophyllum obscurum Xanthophyllum montanum Xanthophyllum ovatifolium Xanthophyllum malayanum Xanthophyllum macrophyllum Xanthophyllum longum Xanthophyllum lineare Xanthophyllum lateriflorum Xanthophyllum purpureum Xanthophyllum pulchrum Xanthophyllum pauciflorum Xanthophyllum parvifolium Xanthophyllum papuanum Xanthophyllum palawanense Xanthophyllum pachycarpon Xanthophyllum eurhynchum Xanthophyllum colubrinum Xanthophyllum cockburnii Xanthophyllum cochinchinense Xanthophyllum ceraceifolium Xanthophyllum burkillii Xanthophyllum brevipes Xanthophyllum bombayanum Xanthophyllum celebicum Xanthophyllum ancolanum Xanthophyllum sylvestre Xanthophyllum vitellinum Xanthophyllum velutinum Xanthophyllum novoguinense Xanthophyllum nitidum Xanthophyllum nigricans Xanthophyllum ngii Xanthophyllum monticola Xanthophyllum rectum Xanthophyllum penibukanense Xanthophyllum punctatum Xanthophyllum pubescens Xanthophyllum poilanei Xanthophyllum octandrum Xanthophyllum petiolatum Xanthophyllum ferrugineum Xanthophyllum flavescens Xanthophyllum excelsum Xanthophyllum havilandii Xanthophyllum inflatum Xanthophyllum incertum Xanthophyllum impressum Xanthophyllum lanceatum Xanthophyllum contractum Xanthophyllum borneense Xanthophyllum bicolor Xanthophyllum bibracteatum Xanthophyllum wrayi Xanthophyllum sulphureum Xanthophyllum subcoriaceum Xanthophyllum trichocladum Xanthophyllum reticulatum Xanthophyllum venosum Xanthophyllum neglectum Xanthophyllum pseudoadenotus Xanthophyllum pedicellatum Xanthophyllum ramiflorum Xanthophyllum ellipticum Xanthophyllum heterophyllum Xanthophyllum angustigemma Xanthophyllum korthalsianum Xanthophyllum crassum Xanthophyllum chartaceum Xanthophyllum brigittae Xanthophyllum brachystachyum Xanthophyllum geminatum Xanthophyllum geesinkii Xanthophyllum fragrans Xanthophyllum eglandulosum Xanthophyllum eberhardtii Xanthophyllum ionanthum Xanthophyllum griffithii Xanthophyllum andamanicum Xanthophyllum amoenum Xanthophyllum adenotus Xanthophyllum discolor Xanthophyllum clovis Xanthophyllum bullatum Xanthophyllum cucullatum Xanthophyllum albicaule Xanthophyllum oliganthum Xanthophyllum yunnanense Xanthophyllum hainanense Xanthophyllum laevis Xanthophyllum acuminatissimum Xanthophyllum manickamii Xanthophyllum beccarianum Xanthophyllum philippinense Xanthophyllum erythrostachyum Xanthophyllum bracteatum Xanthophyllum ecarinatum