Rhizome 2-5 mm thick, internodes to 2-4.5 cm long; vascular strands with bundle sheath fully sclerified, rarely 2 cells thick; sclerenchyma strands absent. Rhizome scales peltate, appressed to spreading (mostly), 4.5-6.5 by 1-2 mm, acute, brown, evenly coloured or with a lighter margin, entire to remotely and weakly dentate, rarely short-dentate, often with irregular protuberances. Fronds simple, dimorphic. Fertile fronds with stipe 5.5-24 cm long; lamina 6-21 by 1.7-6 cm, index 2.8-5.2, widest at 0.2-0.4 from base. Sterile fronds with stipe 2-19 cm long; lamina 6-22 by 2-9.5 cm, index 1.7-3.4, widest at 0.2-0.4 from base; base cuneate, apex obtuse, acute, to acuminate/caudate. Venation: main veins on upper surface raised, distinct. Hydathodes frequent, calcareous scales not persistent. Margin cartilaginous, thickened; without notches or notches sporadically present. Sori round to elongate, occasionally confluent across the connecting veins to transverse coenosori, in one row between adjacent main veins (rarely in two rows), in 5 or more rows between costa and margin (rarely less), 3-4 mm wide, superficial to slightly sunken.