Rhizome slender, woody, densely scaly; scales reddish brown, brown, dark brown, or nearly black, rarely whitish, opaque, ovate-lanceolate or lanceolate, margin entire, toothed, or ciliate, apex attenuate or setaceous. Fronds remote, articulate to rhizome, monomorphic or dimorphic with longer and narrower fertile fronds. Stipe usually glabrous, rarely pubescent, scaly at base. Lamina simple, hastately 3-lobed, palmately or pinnately divided, sometimes pinnatisect with adnate pinnae, herbaceous or leathery, glabrous or pubescent, rarely scaly on abaxial side. Lobes usually lanceolate, margin distinctly cartilaginous, entire, notched, or serrate, apex acuminate or obtuse. Lateral veins (in lamina or segments) distinct; veinlets anastomosing to form areoles with included free veinlets. Sori orbicular, in 1 row on either side of costa, less often linear or in many rows, usually superficial, rarely sunken on abaxial surface and raised on adaxial surface. Sporangia with long stalk, annulus with 14 hardened cells. Spores brown, ellipsoid, surface tuberculate or verrucate. x = 12, n = 36, 37.
Main lateral veins distinct; scales not clathrate