Rhizome 1.5-8 mm thick, internodes 1-6 cm long; vascular strands with bundle sheath sclerified, to 2 cells thick; sclerenchyma strands absent to many. Rhizome scales peltate, appressed to spreading, 2-8.5 by 1-2.6 mm, obtuse, acute or contracted to a narrow acumen, apex flat or distinctly cucullate, straw-coloured to brown, evenly coloured, mottled or with a darkened apex, entire to strongly dentate (often coarsely and irregularly). Fronds simple, dimorphic. Fertile fronds with stipe 1.5-23 cm long; lamina 4-28 by 0.8-5.3 cm, index 2.7-10, widest at 0.2-0.5 from base. Sterile fronds with stipe 1-20 cm long; lamina 3-30 by 0.9-10 cm, index 1.6-6.4 widest at 0.2-0.5 from base. Venation: main veins on upper surface not raised, indistinct, or raised, distinct; veinlets free and anastomosing, free veinlets excurrent and recurrent. Hydathodes absent. Margin not or hardly differentiated, without notches or notches sometimes regularly present. Sori round, elongate or confluent across connecting veins to coenosori, in one row between adjacent main veins (rarely two rows, near the costa at the base of the lamina), in 2-many rows between costa and margin, 3-4 mm wide, superficial to slightly sunken.
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