Rhizome 2.5-5 mm thick, internodes to 1-5 cm long; vascular strands with bundle sheath fully sclerified; the sclerenchyma strands very many. Rhizome scales peltate, spreading to (rarely) squarrose, 3-7 by 1-1.5 mm, acute, straw-coloured to brown, evenly coloured (rarely), or with a lighter margin, remotely and weakly to strongly dentate. Fronds simple, dimorphic. Fertile fronds with stipe 2-19 cm long; lamina 3.5-32 by 0.2-0.3(-0.8) cm, linear. Sterile fronds with stipe 0.5-18 cm long; lamina 4.5-20 by 0.7-6.8(-10) cm, index 2.1-6.5, widest at 0.2-0.5 from base; apex rounded to long-acuminate. Venation: main veins on upper surface raised, distinct; veinlets anastomosing, free veinlets excurrent and recurrent. Hydathodes absent. Margin cartilaginous, flat (rarely) or thickened; notches sporadically to regularly present, absent in fertile fronds. Sori in a longitudinal coenosorus (often interrupted), or (rarely, in transitional fronds) transversely elongated, singly between costa and margin, 2-4 mm wide, superficial.