Terrestrial and climbing plants. Rhizome creeping or climbing, stout, terete, meristelic, with 40 or more vascular bundles in 3 circles, scaly only at apex; scales brown to blackish, orbicular, ovate, or ovate-lanceolate, margin uneven, peltate, caducous. Fronds distant, strongly dimorphic, stipe glabrous except when very young; lamina imparipinnate, leathery or stiffly papery, both surfaces glabrous; sterile pinnae broadly lanceolate, articulate to rachis (apical pinna not articulate), shortly stalked, with single gland on base of each costa, margin sharply toothed; veins anastomosing, forming single row of areoles along each side of costa, other veins free; fertile pinnae linear, lamina very reduced, margin entire. Sporangia acrostichoid, covering whole abaxial surface of fertile pinnae. Spores ellipsoid, with prominent tubercles on distal face. x = 37.
Ferns with long scandent rhizomes. Fronds dimorphic. Lamina simply pinnate; sterile laminae with all but terminal pinna articulated at rachis; margins serrate; sterile pinnae oblong-lanceolate; bases unequal, not lobed. Sori continuous on either side of costa, exindusiate. Stem with a few large central and numerous smaller, peripheral vascular bundles.
Rhizome slender, long-creeping or climbing; veins forming one series of narrow areoles by costa (seen near apex of leaflet); fertile fronds normally wholly fertile.