Plants large. Stems creeping to arched-ascending, 2-or 3-pinnate; paraphyllia mostly foliose, lanceolate or filiform-branched; axillary hairs 2-6(-7)-celled. Stem leaves ovate; margins plane, reflexed, or revolute, papillose-serrulate proximally, often serrulate distally by projecting cell ends; costa ending before apex; laminal cells rounded-to oblong-hexagonal, rhombic, or elliptic, coarsely 1-papillose abaxially, papillae sometimes 2-fid, appearing multipapillose. Branch leaves ovate; margins incurved, erect, or recurved, entire to denticulate; apex usually acute; costa ending before apex, not covered with cells apically; laminal cells rounded, hexagonal, elliptic, or oblong-rhombic, 1-papillose. Sexual condition dioicous; perichaetial leaves lanceolate or ovate-lanceolate, margins sometimes ciliate, costa ending near apex, laminal cells smooth or somewhat papillose. Seta smooth or rarely papillose. Capsule inclined to horizontal, curved-cylindric, asymmetric; annulus 2-4-seriate; operculum conic or rostrate from convex-conic base; endostome segments not perforate, cilia in groups of 2-4.
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