Inflorescences unisexual, sometimes bisexual, borne in the leaf axils and/or (especially the pistillate ones) on the older wood down to the trunk, globose to obovoid-capitate, with a thick rhachis and numerous peltate, long stipitate bracts.
Fruits somewhat drupaceous, embedded in the soft middle layer of the infructescence; seed large, with remnants of endosperm; cotyledons unequal, curved, one thick, the other thin.
Staminate flowers: perianth 2–4(5)-lobed; stamens 2–4, straight in bud; pistillode usually absent.
Leaves almost distichous, pinnately veined; stipules fully amplexicaul, free.
Pistillate flowers without a perianth, stigmas 2, filiform, equal.
Trees, dioecious or sometimes monoecious.