Staminate flowers peripheral, without a perianth, initially covered by the membranous expanded margin of the receptacle, (this cover tearing at anthesis, leaving a tubular part around the pistillate flower and a fringe-like marginal pan); stamens without distinct floral arrangement, straight before anthesis.
Fruit forming a drupaceous whole with the enlarged, fleshy, dark purple to red receptacles crowned with the remnants of the stamens and the marginal part of the receptacle.
Pistillate flower 1, central, partly adnate to the receptacle; perianth 4-lobed; ovary adnate to the perianth, stigmas 2, ligulate, equal.
Inflorescences in the leaf axils, bisexual, initially enveloped by 2 coriaceous bud-scales, involucrate.
Leaves distichous, pinnately veined (to subtriplinerved); stipules fully amplexicaul, connate.
Seed large, without endosperm; cotyledons equal, thick and fused.
Trees, monoecious.