Fruit of 5 samara-like, single-seeded carpels each with a unilaterally produced wing.
Ovary of 5 coherent carpels and with 5 coherent styles; ovules c. 12, 2-seriate.
Petals 5, with a contorted aestivation, tomentose, with a basal claw.
Stamens many; staminodes 5, ovate, with a contorted aestivation.
Trees with petiolate, palmately lobed leaves; stipules present.
Calyx campanulate with 5 valvate lobes.
Androgynophore conspicuous.