Typha L.

Cattail (en), Massette (fr)

Genus

Angiosperms > Poales > Typhaceae

Characteristics

Plants of fresh to slightly brackish wetlands, often emergent. Rhizomes at base of erect shoots, mostly horizontal, unbranched, to 70 cm  5--40 mm, starchy, firm, scaly. Erect shoots vegetative or flowering, single at rhizome apices or arising from shoot bases, thus clustered, unbranched, to 4 m, elliptic in cross section; stems often somewhat compressed distally, aerenchyma absent. Foliage leaves persistent, intergrading proximally with scale leaves, to 15 on each flowering shoot; blade twisted into loose helix, mostly slightly oblanceolate, thickly concave-convex or plano-convex proximally to thinly plane distally (abaxially keeled in the Old World Typha. eleiphantina Roxburgh); mucilage-secreting glands numerous in adaxial surface of sheath and sometimes proximally on blade, colorless to brown, roughly rectangular. Inflorescences: staminate scales shorter than or exceeding flowers; pistillate spikes usually persisting into winter, when dry fruiting flowers often falling in masses; pistillate bracteoles absent or numerous, colorless except for brown apical blade at spike surface, filiform, blade club-shaped to lanceoloid. Staminate flowers: anthers dehiscing longitudinally, 4-sporangiate. Pistillate flowers: pistil hairs colorless and wholly filiform, or apically enlarged and brown, exceeded by stigmas; carpodia obovoid, spongy, bearing rudimentary styles. x = 15.The extensive literature on Typha has been reviewed (C. M. Finlayson et al. 1983; J. B. Grace and J. S. Harrison 1986; J. W. Thieret and J. O. Luken 1996). A modern taxonomic revision is much needed, especially for eastern Asia, adjacent islands, and South America (S. G. Smith 1987); the latest worldwide monograph is that by E. M. Kronfeld (1889). The center of diversity (ca. 6 species) is central Eurasia.
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Perennial, palustrial or aquatic herbs with a creeping rhizome; stems erect, solid, submerged at the base. Leaves biseriate, partly radical or subradical, partly cauline, lower congested, higher remote, elongate-linear, rather thick and spongy, blunt-margined; their sheathing bases excreting slime on their inner side. Flowers very numerous, very closely packed in 2 or less often 3, superposed, contiguous or more or less remote terete unisexual spikes; upper spike male; the 1-2 lower ♀; all spikes at the base with a foliaceous bract which falls off long before anthesis; the ♀ spikes here and there between the flowers often with a similar bract. ♂ Flowers consisting of 3 flat hairs together surrounding 2-5 stamens; anthers basifixed, linear, 2-celled; connective shortly produced; cells back to back, bursting longitudinally; pollen-grains free or cohering in tetrads. Rachis of ♀ spathe closely studded with patent cylindrical thickish excrescences; between these excrescences and on their basal part beset with flowers containing a fertile ovary; higher part of the excrescences bearing rudimentary ovaries. ♀ Flowers with or without a very narrow bracteole; bracteole with a more or less broadened, often dentate-acuminate apex either entirely hidden by the flowers or their apex visible externally. Ovary borne by a long very thin stalk (gynophore) which bears long hairs on its base, fusiform, 1-celled; style distinct thin; stigma broadened, unilateral, linear or spathulate. Fruit small, fusiform, or elongate-ovoid, falling off together with its stalk from the pilose axis of the spike, finally bursting by a longitudinal slit; seed pendulous, striate; endosperm mealy; embryo narrow, straight, nearly as long as the seed.
Stout glabrous marsh herbs with sympodial branched rhizomes covered by distichous cataphylls and having terminal erect shoots. Lower leaves transitional, short-bladed; upper leaves with a long sheath closely enveloping the stem, often auricled at the junction with the blade, and a linear obtuse or acute blade gradually widening from a narrow subpetiolar base, semi-cylindric to lenticular in section, sometimes keeled beneath. Male inflorescence apical, dense; flowers reduced to a pedicellate group of 2–4(–7) stamens with the filaments fused below; anthers dithecous, oblong, with the connective produced above; bracteoles filiform, spathulate or spathulate-laciniate; pollen grains shed singly or in tetrads. Female inflorescence a single spike beneath the ♂, remote from it or contiguous, rarely with 1 or 2 additional ♀ spikes remote from each other beneath the first; flowers with the perigonous hairs and bracteoles of similar length, the styles conspicuously longer; bracteoles filiform, broadening into an expanded ovate or spathulate lamina; carpodia about as long as the bracteoles. Follicle narrowly ellipsoid; seed fusiform.
Herbs, perennial, in marshes or aquatic, with creeping rhizomes. Leaves alternate, erect, distichous, linear, usually spongy, margin entire, sheathed at base. Flowers unisexual, minute, numerous, densely crowded in a cylindric spike with lower part female and upper part male; bracts leaflike. Perianth absent. Male flowers consisting of 1-3 stamens usually connate at base of filaments, surrounded by hairs; anthers 2-thecous, basifixed, longitudinally dehiscent; filaments short; pollen grains in monads or tetrads. Female flowers: ovary 1-loculed, on a long capillary stalk with many fine hairs or bracteoles at base; styles capillary; stigmas broadened or spatulate; ovule 1; sterile ovary without style. Fruit minute, falling off together with stalk.
Massive semiaquatic perennial herbs with extensive fleshy rhizomes. Leaves elongate, flat, equitant. Flowering stems elongate, terete; flowers monoecious, very numerous in two adjacent dense, cylindric spikes, the staminate above the pistillate, each usually subtended by a reduced, spathe-like leaf; perianth reduced to bristles or hairs; ovary stipitate, 1-2-celled.
Characters of the family. 10, cosmop.
Characters as for the family.
See Typhaceae.
Life form perennial
Growth form herb
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Foliage retention deciduous
Sexuality monoecy
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Root system rhizome
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Environment

Marshy places, shallow pools of fresh or brackish water, often gregarious.
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Hardiness (USDA) 5-12

Usage

Uses. The rhizomes which are rich in starch are eaten in many regions where food is scarce, or in periods of famine. The leaves are used for thatching huts, for matting and for coarse basket-work. The spikes are often used for decoration; the plush of the ripe ♀ spadices was formerly used for stuffing pillows.
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Cultivation

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