Utricularia scandens Benj.

Species

Angiosperms > Lamiales > Lentibulariaceae > Utricularia

Characteristics

Small or sometimes very small, probably annual terrestrial. Rhizoids numerous, capillary, a few mm to a few cm long, c. 0.1 mm thick, bearing simple papillose branches up to 1.5 mm long. Stolons often numerous, capillary, sparsely branched, up to several cm long, c. 0.15 mm thick. Leaves few to many, petiolate, the lamina narrowly linear, with apex rounded or subacute, 0.5-1 mm wide, 1-nerved; total length up to 2 cm. Traps moderately numerous on the rhizoids, stolons and leaves, globose, shortly stalked, 0.5-1 mm long, the mouth basal with 2 dorsal, simple, subulate, glandular appendages and a single short to relatively long, truncate or apically ± bifid, glandular ventral appendage. Inflorescence erect or twining, solitary, simple, 3-35 cm long; peduncle somewhat angular, glabrous throughout or papillose below, 0.2-0.6 mm thick. Scales moderately numerous, similar to the bracts. Bracts basifixed, broadly ovate-deltoid, with apex acute or acuminate, 1-1.5 mm long. Bracteoles basifixed, narrowly linear or subulate, with apex acute, as long as or somewhat shorter than the bracts. Flowers 1-8, the raceme elongate and often bearing, on its axis, bracts subtending aborted or dormant buds alternating with those which subtend flowers; pedicels erect, filiform, relatively broadly winged, up to 1.5 cm long and 1 to 5 times as long as the flowering calyx. Calyx lobes subequal, the lower lobe slightly smaller, ovate, the upper lobe with apex acute or acuminate, the lower lobe with apex shortly and acutely bifid, 2-4 mm long at anthesis, up to 6 mm long in fruit. Corolla yellow, 0.5-1.5 cm long; upper lip constricted below the middle, the superior part oblong to transversely elliptic, with apex rounded or emarginate, the inferior part quadrate; lower lip limb circular or broadly obovate, the base distinctly swollen, with 2 or 4 rounded, longitudinal ridges, the apex rounded or obscurely 2-3 crenate; palate papillose; spur subulate, with apex acute, curved or straight, longer than and widely diverging from the lower lip. Filaments straight, c. 1 mm long, the anther thecae distinct. Ovary ovoid; style short; stigma lower lip semicircular, the upper lip similar but smaller. Capsule ovoid, dorsiventrally compressed, 2-2.5 mm long, the wall membranous, dehiscing by a single, ventral, longitudinal, marginally thickened slit. Seeds obliquely obovoid, c. 0.2 mm long, the testa cells elongate, with the anticlinal boundaries raised, straight or slightly sinuate, microscopically tuberculate, the periclinal walls concave, microscopically tuberculate. Collected in flower more or less throughout the year but probably mostly during and immediately after the wet season.
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Terrestrial. Rhizoids numerous, capillary, with papillose branches c. 0.5 mm long. Stolons few capillary branched. Foliar organs on the stolons, linear, membranous, up to 1 cm long, c. 1 mm wide, 1-nerved, apex rounded or subacute. Traps on the vegetative organs, globose, 0.6-1 mm long, shortly stalked, mouth basal, upper lip with 2 simple subulate recurved appendages, lower lip with 1 shorter truncate or shortly bifid usually apically glandular appendage. Inflorescence twining, 3-35 cm long; peduncle filiform, terete, glabrous; flowers 1-8, distant, with 1 or 2 sterile bracts on the internodes; scales few, similar to the bracts; bracts broadly ovate-deltoid, acute or acuminate, 1-1.5 mm long, nerveless; bracteoles linear-lanceolate, about as long as the bract, nerveless; pedicels erect, filiform, narrowly winged, about as long as the flowering calyx. Calyx lobes subequal, ovate or broadly ovate, 2.5-3 mm long at anthesis, up to 5 mm long in fruit, upper slightly larger, apex shortly acuminate, lower with apex shortly bi-or tridentate. Corolla yellow, 5-10 mm long, upper lip oblong, usually shorter and narrower than upper calyx lobe, apex rounded, entire or emarginate or ± bifid, lower lip larger, ± orbicular, apex rounded, entire or 2-3-crenate, palate ± raised, smooth or 2-4-gibbous, spur subulate, acute, curved, longer than and widely diverging from the lower lip. Filaments linear, ± straight, anther thecae ± confluent. Ovary ovoid, style short, indistinct, stigma lower lip semi-orbicular, upper lip similar, smaller. Capsule oblong-ovoid, dorsiventrally compressed, membranous, 2-2.5 mm long, dehiscing by a single ventral longitudinal marginally thickened slit. Seeds numerous, broadly ellipsoid or ovoid, c. 0.2 mm long, testa thin, smooth, reticulate, reticulations elongate, hilum lateral, prominent.
Terrestrial herb.. Rhizoids and stolons capillary, few from the base of the scape.. Leaves few, branch-opposed on the stolons, linear, 1-nerved, up to 10 mm. long and 1 mm. wide.. Traps scattered on the stolons and leaves, globose, shortly stalked, glandular, 0.6–1.0 mm. long; mouth basal; upper lip with 2 simple subulate appendages; lower lip with a single shorter truncate or shortly bifid appendage.. Inflorescence erect or twining, 3–35 cm. high; scape filiform, glabrous; flowers 1–8, distant, usually alternating with sterile bracts; scales few, similar to the bracts; bracts basifixed, broadly ovate-deltoid, acute or acuminate, nerveless, 1–1.5 mm. long; bracteoles linear-lanceolate, ± 1 mm. long; pedicels spreading or erect, 1–3 times as long as the flowering calyx, narrowly winged.. Calyx-lobes broadly ovate, 2.5–3 mm. long at anthesis, up to 5 mm. long in fruit, decurrent, slightly unequal; upper longer, acute or acuminate; lower shortly 2–3-dentate.. Corolla yellow, 5–15 mm. long; upper lip shorter than to about twice as long as the upper calyx-lobe, oblong to orbicular, apex ± emarginate; lower lip orbicular, apex rounded, entire or 2–3-crenate; palate raised, usually 2–4-gibbous; spur subulate, acute, ± curved.. Filaments linear, ± straight; anther-thecae ± confluent.. Ovary ovoid; style short; stigma-lips semi-orbicular, the upper smaller.. Capsule oblong-ovoid, dorsiventrally compressed, 2–2.5 mm. long, dehiscing by a longitudinal abaxial slit with thickened margins.. Seeds ovoid or elliptic, smooth, ± 0.2 mm. long; hilum prominent; testa-cells distinct, considerably longer than broad.
Annuals, terrestrial. Rhizoids and stolons capillary, branched. Traps on rhizoids, stolons, and leaves, stalked, globose, 0.5-1 mm, mouth basal; appendages 2, dorsal, simple, subulate, glandular. Leaves few to many, glabrous; leaf blade narrowly linear, 5-10 × 0.4-1 mm, membranous, vein 1, base attenuate onto petiole, margin entire, apex rounded to subacute. Inflorescences erect or twining, 1.5-35 cm, (1-)3-8-flowered, glabrous; peduncle terete but somewhat angular, 0.2-0.6 mm thick; scales few to numerous, similar to bracts; bracts basifixed, broadly ovate-deltoid, 0.5-1.5 mm, apex acute to acuminate. Pedicel suberect, 2.5-3 mm, filiform, winged; bracteoles basifixed, narrowly linear to subulate, as long as or slightly shorter than bracts, apex acute. Calyx lobes ovate to elliptic, 2-3 mm at anthesis, to 5 mm in fruit; lower lobe subequal to or slightly smaller than upper lobe, apex shortly 2-cleft; upper lobe apex acute to acuminate. Corolla yellow, 5-11 mm; lower lip suborbicular, base distinctly swollen, apex rounded to obscurely crenate; spur subulate, longer than lower corolla lip, widely divergent, curved or straight, apex acute; palate papillose; upper lip oblong, apex rounded to emarginate. Filaments ca. 1 mm, straight; anther thecae distinct. Ovary ovoid, dorsiventrally compressed; style short; stigma lower and upper lips semicircular. Capsule ovoid, 2-2.5 mm, dorsiventrally compressed, dehiscing by a longitudinal ventral slit. Seeds obliquely obovoid, 0.2-0.3 mm; seed coat with prominent elongate reticulations. Fl. Jun-Sep, fr. Jul-Oct.
Inflorescence erect or twining, 3–35 cm. high; peduncle filiform, glabrous; flowers 1–8, distant, usually alternating with sterile bracts; scales few, similar to the bracts; bracts basifixed, broadly ovate-deltoid, acute or acuminate, nerveless, 1–1.5 mm. long; bracteoles linear-lanceolate, more or less 1 mm. long; pedicels spreading or erect, 1–3 times as long as the flowering calyx, narrowly winged.
Corolla yellow, 5–15 mm. long; superior lip shorter than to about twice as long as the upper calyx lobe, oblong to circular, with apex more or less emarginate; inferior lip circular, with apex rounded, entire or 2–3-crenate; palate raised, usually 2–4 gibbous; spur subulate, acute, more or less curved.
Traps scattered on the stolons and leaves, globose, shortly stalked, glandular, 0.6–1.0 mm. long; mouth basal, with 2 dorsal simple subulate appendages, a single shorter truncate or shortly bifid ventral appendage.
A widespread species, subsp. scandens occurring throughout E. & S. tropical Africa, Madagascar and tropical Asia, and the following subspecies (Utricularia scandens subsp. schweinfurthii) in W. Africa.
Calyx lobes broadly ovate, 2.5–3 mm. long at anthesis, up to 5 mm. long in fruit, deccurent, slightly unequal; upper longer, with apex acute or acuminate; lower with apex shortly 2–3-dentate.
Seeds ovoid or elliptic, smooth, more or less 0.2 mm. long; hilum prominent; testa-cells distinct, considerably longer than broad.
Capsule oblong-ovoid, dorsiventrally compressed, 2–2.5 mm. long, dehiscing by a longitudinal ventral slit with thickened margins.
Leaves few, branch-opposed on the stolons, linear, 1-nerved, up to 10 ×1 mm.
Ovary ovoid; style short; stigma-lips semi-circular, the superior smaller.
Filaments linear, more or less straight; anther-theacae confluent.
Rhizoids and stolons capillary, few from the base of the peduncle.
Terrestrial herb.
Life form annual
Growth form herb
Growth support climber
Foliage retention -
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.03 - 0.35
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway -

Environment

Damp places twining up grasses, in Malaya at 1200 m, ascending in New Guinea from the lowland to 2700 m. Fl. April-Aug.
Light -
Soil humidity 7-9
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 8-12

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Utricularia scandens unspecified picture

Distribution

Utricularia scandens world distribution map, present in Australia, China, India, Iceland, Madagascar, Papua New Guinea, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:527085-1
WFO ID wfo-0000417444
COL ID 7F575
BDTFX ID -
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Utricularia capillacea Utricularia scandens Utricularia wallichii Polypompholyx madecassa Utricularia gibbsiae Utricularia macrolepis Utricularia scandens subsp. scandens Utricularia bifida var. wallichiana Utricularia scandens var. scandens

Lower taxons

Utricularia scandens subsp. schweinfurthii