Utricularia L.

Bladderworts (en), Utriculaire (fr)

Genus

Angiosperms > Lamiales > Lentibulariaceae

Characteristics

Herbs, annual or perennial, terrestrial, epiphytic or aquatic. Vegetative parts not clearly differentiated but consisting of stems modified to function as roots, stems, leaves and specialized organs (traps) for the capture of small organisms. Root-like organs (rhizoids) usually descending from the base of the inflorescence, usually filiform. Stem-like organs (stolons) arising with the rhizoids at the inflorescence base; in the terrestrial and epiphytic species usually short and delicate but sometimes developing into fleshy tubers; in the aquatic species usually more robust and longer. Foliar organs (leaves) either rosulate at the inflorescence base or alternate, opposite or verticillate on the stolons; in the terrestrial and epiphytic species entire, erect or thalloid, capillary, linear, orbicular or peltate; in the aquatic species ± dichotomously divided into capillary segments. Traps hollow, globose or ovoid, usually stalked, with a mouth which may be basal (adjacent to the stalk), lateral or terminal (opposite to the stalk); mouth provided externally with 2 lips which bear very diverse appendages. Inflorescence racemose, bracteate; scape usually simple, sometimes branched, usually filiform, erect or twining, usually glabrous, sometimes papillose, glandular or hairy, provided (especially in the terrestrial and epiphytic species) with sterile bracts (scales); raceme usually elongated, rarely short and subcapitate; pedicels usually short, terete, flattened or ± winged, often reflexed or recurved in fruit; bracts persistent, basifixed or produced below the point of insertion or peltate; bracteoles 2 or absent or sometimes ± fused with the bract, inserted with the bract at the base of the pedicel or rarely with the calyx-lobes at the apex of the pedicel. Calyx-lobes 2, ± equal or sometimes very unequal, usually free, sometimes ± united at the base, persistent and usually accrescent, sometimes very markedly so; upper lobe usually entire, lower lobe usually emarginate or bidentate, rarely both lobes dentate or fimbriate. Corolla bilabiate, glabrous, glandular or pubescent; throat closed or sometimes open; upper lip usually ± erect, limb entire, emarginate or bilobed; lower lip usually larger, spurred or rarely saccate at the base, palate usually raised and gibbous, limb spreading or deflexed, entire, emarginate or ± deeply 2–5-lobed; spur ± parallel to the lower lip or divergent at an acute or obtuse angle or rarely in the same plane. Stamens 2, inserted at the base of the corolla; filaments straight or curved, usually twisted, sometimes winged; anthers dorsifixed, ± ellipsoid, the thecae ± confluent. Ovary globose or ovoid, unilocular; style usually short, often indistinct, persistent; stigma bilabiate, lower lip usually much larger than the upper which may be obscure or obsolete; ovules 2-many, sessile on a ± fleshy basal or free central placenta, anatropous. Fruit a capsule, ± globose, dehiscing by longitudinal slits or by pores or circumscissile or indehiscent. Seeds 1–many, usually small or very small, globose, ovoid, truncate conical, narrowly cylindrical, fusiform, lenticular or prismatic, smooth, verrucose, reticulate, glochidiate or variously winged. Embryo undifferentiated.
More
Annual or perennial aquatic terrestrial or epiphytic herbs always of damp places, without true roots or leaves but with stems modified in various ways to function as rhizoids, stolons and foliar organs, all species bearing small complex bladder-like traps for the capture and digestion of small aquatic organisms. Inflorescence racemose, peduncled, usually simple, bracteate; sterile bracts (scales) often present on the peduncle and sometimes also on the inflorescence axis; two bracteoles often present, almost always at the base of the pedicel, usually free, rarely ± connate with the bract. Bracts very varied, basifixed, medifixed or variously produced below the point of attachment. Calyx 2-lobed, usually ± accrescent, the lobes ± equal or variously dissimilar, usually free, sometimes ± connate at the base. Corolla bilabiate, yellow, various shades of violet or purple, white or rarely blue or red; upper lip entire or 2-or more-lobed; lower lip with an entire or 2-5-lobed limb, a ± raised, often gibbous palate and a usually subulate or conical spur, in a few species reduced to a short sac. Stamens 2 inserted at the base of the upper lip; filaments usually short, linear, often curved and often ± flattened and dilated above; anthers dorsifixed, ± ellipsoid, thecae ± confluent. Ovary globose or ovoid, ovules 2-many on a free basal or free central ± fleshy placenta; style usually short; stigma bilabiate, the lower lip usually much larger. Capsule globose or ovoid, dehiscing very variously by longitudinal slits, dorsiventral or rarely lateral valves, pores or circumscissile or rarely indehiscent. Seeds 1-many, very variously shaped and sculptured.
Herbs, annual or perennial, aquatic, terrestrial or epiphytic, always of damp places, without true roots or leaves but with stems modified in various ways to function as tubers, rhizoids, stolons, and + leaflike photosynthetic organs, all species bearing small complex bladderlike traps for the capture and digestion of small organisms. Inflorescences pedunculate, racemose, usually simple, brac-teate, sterile bracts (scales) often present on the peduncle and sometimes also on the inflorescence axis; bracts varied, basifixed, medifixed or variously produced below the point of insertion; bracteoles 2 or absent, rarely connate with the bract. Flowers with the calyx 2-lobed, usually accrescent; corolla usually spurred, yel-low or various shades of violet or purple, or white, rarely red, the upper lip en-tire or + 2-lobed, the lower lip entire or 2-5-lobed; stamens usually short, the filaments often winged, the anthers globose or ovoid; ovary ? globose, the ovules 2-many. Capsules globose to ovoid, dehiscing variously by longitudinal slits, dorsiventral or lateral valves or pores, circumscissile or indehiscent; seeds 1-many, variously shaped.
Herbs, perennial or annual, terrestrial, epiphytic, or aquatic, without true roots. Stems modified into rhizoids and stolons, rarely developed. Traps on rhizoids, stolons, and/or leaves, small, bladderlike. Leaves alternate or in a basal rosette, simple to many × divided, veins 1-3, unbranched, dichotomously branched, or pinnately branched. Inflorescences racemose or flowers solitary, pedunculate, usually simple, seldom branched, erect to twining, bracteate; bracts and bracteole often present, scalelike, sometimes basisolute (with base extending below point of insertion). Calyx parted from base into 2 equal or unequal lobes, lobes sometimes apically 2-parted. Corolla lower lip larger than upper lip; lower lip entire or 2-or 3(-6)-lobed, spurred, palate variously raised; upper lip entire or 2-or 3-lobed. Anther thecae confluent or distinct. Capsule adaxially loculicidal, both abaxially and adaxially loculicidal, or circumscissile, rarely indehiscent. Seeds few, many, or rarely 1 per capsule, variously appendaged.
Annual or perennial, aquatic, terrestrial or epiphytic herbs. Roots 0, but root-like structures present. Stems modified to form rhizoids, stolons and leafy shoots. Lvs usually small, entire when terrestrial or emergent, capillary and much-divided when submerged, all with small bladder-like traps. Infl. racemose, emergent in aquatic spp., usually with a simple scape or peduncle. Fls 1-several. Calyx 2-lipped, divided almost to base, usually accrescent. Corolla yellow, white or purple; upper lip entire or lobed, mainly small; lower lip entire or 2-5-lobed, generally larger than upper lip, with palate or pouch present at base, ± raised and often gibbous; spur usually subulate or conic. Filaments usually curved, often dilated and flattened; anthers convergent. Stigma unequally lobed. Ovules 1-numerous. Capsule globose or ovoid, dehiscing in various ways. Seed very variable in shape and sculpturing.
Inflorescence racemose, (but sometimes with a single flower) bracteate; peduncle usually simple, sometimes branched, usually filiform, erect or twining, usually glabrous, sometimes papillose, glandular or hairy, often provided (especially in the terrestrial species) with sterile bracts (scales); raceme usually elongated, rarely short and subcapitate; pedicels usually short, terete, flattened or more or less winged, often deflexed or decurved in fruit; bracts persistent, basifixed or produced below the point of insertion or peltate; bracteoles 2 or absent or sometimes more or less fused with the bract, inserted with the bract at the base of the pedicel or rarely with the calyx lobes at the apex of the pedicel.
Herbs, terrestrial or aquatic, affixed or freely suspended in water; stolons usually present (absent only in a few species, all Australian). Traps bladder-like, variously inserted on the vegetative parts, usually stalked, globose or ovoid; mouth basal, lateral or terminal, usually provided with appendages of very various kinds and closed by a door which bears on its outer surface varied glands or hairs. Inflorescence racemose or by reduction 1-flowered; peduncle often bearing sterile bracts (scales) between its base and the lowermost flower. Calyx lobes 2 or rarely 4.
Cal parted to the base into upper and lower segments, the upper often wider; cor bilabiate, the upper lip usually subentire or shallowly 2-lobed, the lower entire or 3-lobed, usually elevated at the base into a prominent palate, the tube prolonged at base into a spur or sac; herbs of water or wet soil, the lvs (lf-like branches?) linear and entire or variously dissected, in most spp. provided with small bladders that catch tiny aquatic animals; fls yellow or violet, in short racemes (or apparently solitary and terminal), each subtended by a small bract. 150, cosmop.
Scapes with 1-several fls; calyx deeply 2-lobed, segs often enlarging as capsule matures. Corolla spurred at back, 2-lipped; upper lip erect, entire or 2-lobed; lower lip spreading, us. 3-6-lobed; palate us. well-developed; stamens 2, ± curved; style short; stigma 2-lobed. Capsule subglobose, 2-valved, or dehiscing irregularly; seeds ∞. Herbs with filiform lvs and ∞ to few bladders that trap small animals. Widespread genus with some 250 spp. U. monanthos occurs also in Tasmania, and U. lateriflora in Tasmania and Australia. The other N.Z. spp. appear to be endemic.
Corolla bilabiate, glabrous, glandular or pubescent; throat closed or sometimes open; superior lip usually more or less erect, limb entire, emarginate or bilobed; inferior lip usually larger, spurred or rarely saccate at the base, palate usually raised and gibbous, limb spreading or deflexed, entire, emarginate or more or less deeply 2–5-lobed; spur more or less parallel to the inferior lip or divergent at an acute or obtuse angle or rarely in the same plane.
Calyx lobes 2, or rarely 4 in 2 decussate pairs, more or less equal or sometimes very unequal, usually free, sometimes more or less united at the base, persistent and usually accrescent, sometimes very markedly so; upper lobe usually entire, inferior lobe usually emarginate or bidentate, rarely both lobes dentate or fimbriate.
Foliar organs (leaves) either rosulate at the inflorescence base or alternate, opposite or verticillate on the stolons; in the terrestrial and epiphytic species mostly entire, erect or thalloid, capillary, linear, circular or peltate; in the aquatic species usually more or less dichotomously divided into capillary segments.
Traps rosulate, or lateral on the rhizoids, stolons or leaves or rarely terminal on the leaves, hollow, globose or ovoid, usually stalked, with a mouth which may be basal (adjacent to the stalk), lateral or terminal (opposite to the stalk); mouth usually provided externally with very diverse appendages.
Ovary globose or ovoid, unilocular; style usually short, often indistinct, persistent; stigma bilabiate, inferior lip usually much larger than the upper which may be obscure or obsolete; ovules 2-many, sessile on a more or less fleshy free basal placenta, anatropous.
Stem-like organs (stolons) arising with the rhizoids at the inflorescence base; in the terrestrial and epiphytic species usually short and delicate but sometimes developing into fleshy tubers; in the aquatic species usually more robust and longer.
Seeds 1-many, usually small or very small, globose, ovoid, truncate conical, narrowly cylindrical, fusiform, lenticular or prismatic, smooth, verrucose, reticulate, glochidiate, papillose or comose or variously winged.
Stamens 2, inserted at the base of the corolla; filaments straight or curved, usually twisted, sometimes winged; anthers dorsifixed, more or less ellipsoid, the thecae usually more or less confluent.
Vegetative parts not clearly differentiated but consisting of stems modified to function as roots, stems, leaves and specialized organs (traps) for the capture of small organisms.
Fruit a capsule, more or less globose or ovoid, dehiscing by longitudinal slits or by pores or circumscissile or indehiscent.
Root-like organs (rhizoids) usually descending from the base of the inflorescence, usually filiform.
Herbs, annual or perennial, terrestrial, epiphytic or aquatic.
Life form
Growth form herb
Growth support -
Foliage retention -
Sexuality hermaphrodite
Pollination -
Spread -
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Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color -
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Nitrogen fixer -
Photosynthetic pathway -

Environment

Marshes, wet grassfields, swamps, swamp-forest, streams and rivers and open damp sandy ground, a few species epiphytic among moss on trees (and rocks), occurring in Malesia from sea-level to 3660 m.
Light -
Soil humidity -
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Hardiness (USDA) 8-12

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
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Germination treatment -
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Images

Utricularia unspecified picture
Utricularia unspecified picture

Distribution

Utricularia world distribution map, present in American Samoa, Australia, China, Indonesia, India, New Zealand, Panama, Philippines, Papua New Guinea, Thailand, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:30001688-2
WFO ID wfo-4000039907
COL ID 85JD
BDTFX ID 87372
INPN ID 198838
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Lepiactis Biovularia Enskide Cosmiza Askofake Bucranion Calpidisca Diurospermum Hamulia Plectoma Polypompholyx Vesiculina Meionula Utricularia

Lower taxons

Utricularia foliosa Utricularia fimbriata Utricularia firmula Utricularia furcellata Utricularia garrettii Utricularia juncea Utricularia malabarica Utricularia macrorhiza Utricularia janarthanamii Utricularia jamesoniana Utricularia jackii Utricularia inflexa Utricularia humboldtii Utricularia hydrocarpa Utricularia huntii Utricularia bisquamata Utricularia bracteata Utricularia alpina Utricularia beaugleholei Utricularia arenaria Utricularia cymbantha Utricularia unifolia Utricularia tortilis Utricularia tricolor Utricularia warmingii Utricularia pentadactyla Utricularia pubescens Utricularia praetermissa Utricularia mirabilis Utricularia microcalyx Utricularia micropetala Utricularia ochroleuca Utricularia spruceana Utricularia simulans Utricularia tenuissima Utricularia subramanyamii Utricularia subulata Utricularia stygia Utricularia scandens Utricularia schultesii Utricularia babui Utricularia simmonsii Utricularia chiribiquetensis Utricularia erectiflora Utricularia livida Utricularia heterochroma Utricularia hispida Utricularia guyanensis Utricularia campbelliana Utricularia breviscapa Utricularia buntingiana Utricularia bremii Utricularia adpressa Utricularia australis Utricularia aureomaculata Utricularia amethystina Utricularia aurea Utricularia andongensis Utricularia cucullata Utricularia cornuta Utricularia naviculata Utricularia naikii Utricularia physoceras Utricularia praelonga Utricularia prehensilis Utricularia olivacea Utricularia oliveriana Utricularia stellaris Utricularia sandwithii Utricularia benjaminiana Utricularia calycifida Utricularia appendiculata Utricularia nana Utricularia myriocista Utricularia neottioides Utricularia nervosa Utricularia welwitschii Utricularia pusilla Utricularia spiralis Utricularia petertaylorii Utricularia paulineae Utricularia striatula Utricularia reflexa Utricularia quelchii Utricularia steyermarkii Utricularia cornigera Utricularia chiakiana Utricularia inflata Utricularia densiflora Utricularia inthanonensis Utricularia spinomarginata Utricularia phusoidaoensis Utricularia cochleata Utricularia rostrata Utricularia catolesensis Utricularia regia Utricularia resupinata Utricularia reniformis Utricularia geminiscapa Utricularia purpurea Utricularia costata Utricularia tenella Utricularia multifida Utricularia monanthos Utricularia tridactyla Utricularia singeriana Utricularia fistulosa Utricularia tubulata Utricularia lasiocaulis Utricularia kamienskii Utricularia kenneallyi Utricularia leptorhyncha Utricularia choristotheca Utricularia troupinii Utricularia purpureocaerulea Utricularia flaccida Utricularia laciniata Utricularia longifolia Utricularia floridana Utricularia inflata Utricularia radiata Utricularia endresii Utricularia lloydii Utricularia lloydii Utricularia sandersonii Utricularia westonii Utricularia menziesii Utricularia volubilis Utricularia helix Utricularia benthamii Utricularia violacea Utricularia terrae-reginae Utricularia novae-zelandiae Utricularia uniflora Utricularia kimberleyensis Utricularia georgei Utricularia hamiltonii Utricularia triflora Utricularia arnhemica Utricularia albiflora Utricularia asplundii Utricularia odontosepala Utricularia hintonii Utricularia panamensis Utricularia platensis Utricularia bifida Utricularia graminifolia Utricularia minutissima Utricularia caerulea Utricularia salwinensis Utricularia brachiata Utricularia limosa Utricularia punctata Utricularia hirta Utricularia striata Utricularia mannii Utricularia pobeguinii Utricularia stanfieldii Utricularia uliginosa Utricularia quinquedentata Utricularia holtzei Utricularia rhododactylos Utricularia cheiranthos Utricularia capilliflora Utricularia dunlopii Utricularia dunstaniae Utricularia antennifera Utricularia geoffrayi Utricularia lateriflora Utricularia delicatula Utricularia simplex Utricularia warburgii Utricularia podadena Utricularia odorata Utricularia involvens Utricularia delphinoides Utricularia arcuata Utricularia pierrei Utricularia circumvoluta Utricularia recta Utricularia macrocheilos Utricularia heterosepala Utricularia peranomala Utricularia christopheri Utricularia forrestii Utricularia kumaonensis Utricularia steenisii Utricularia pulchra Utricularia moniliformis Utricularia corynephora Utricularia dimorphantha Utricularia uxoris Utricularia mangshanensis Utricularia nelumbifolia Utricularia meyeri Utricularia longiciliata Utricularia perversa Utricularia gibba Utricularia biceps Utricularia blanchetii Utricularia parthenopipes Utricularia tridentata Utricularia nephrophylla Utricularia determannii Utricularia nigrescens Utricularia poconensis Utricularia biovularioides Utricularia julianae Utricularia chrysantha Utricularia fulva Utricularia bosminifera Utricularia cecilii Utricularia praeterita Utricularia albocaerulea Utricularia lazulina Utricularia reticulata Utricularia smithiana Utricularia wightiana Utricularia polygaloides Utricularia foveolata Utricularia vitellina Utricularia letestui Utricularia leptoplectra Utricularia biloba Utricularia muelleri Utricularia jobsonii Utricularia byrneana Utricularia wannanii Utricularia barkeri Utricularia ramosissima Utricularia corneliana Utricularia dichotoma Utricularia linearis Utricularia blackmanii Utricularia petersoniae Utricularia multicaulis Utricularia tetraloba Utricularia raynalii Utricularia inaequalis Utricularia geminiloba Utricularia laxa Utricularia incisa Utricularia rigida Utricularia trichophylla Utricularia triloba Utricularia viscosa Utricularia vulgaris Utricularia minor Utricularia intermedia