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Shrubs or small trees, often unbranched or few branched; at least younger parts ± ferruginous pubescent or tomentose. Leaves often ± crowded towards ends of branches, alternate, usually imparipinnate; petiolar base and rachis joints often constricted (in dried specimens); leaflets 5–13, opposite or subopposite, sessile (usually more so towards the leaf-apex) to shortly petiolulate, entire or toothed. Inflorescence axillary, spiciform or in spiciform panicles, polygamous. Flowers usually in distant glomerules, unisexual or bisexual. Sepals 4–5, united towards the base, imbricate. Petals 4–5, free, imbricate. Stamens 4–5, inserted beneath and between the lobes of the disc, free, unappendaged, reduced in female flowers. Disc fleshy, (3–)4(–5)-lobed. Gynoecium of 4–5 carpels, free or joined only at base, each containing 1 pendulous ovule, absent or vestigial in male flowers; styles 4–5, free. Fruit of drupaceous mericarps developing from 1–4 carpels of each flower. Seed 1 per mericarp.
Shrubs or small trees, monoecious or dioecious, with bitter root cortex and bark. Branches pubescent or puberulent when young. Leaves odd-pinnate; stipules absent; leaflets 3-15; blades ovate or lanceolate, base slightly oblique, margin entire or serrate, apex acuminate. Flowers unisexual, rarely bisexual, in narrow axillary cymose panicles. Sepals 4, small, ovate, oblong-ovate, or deltoid, bases connate, imbricate in bud. Petals 4, small, free, imbricate in bud, oblong-ovate, oblong, or linear. Disk thick, 4-lobed. Stamens 4, only with rudiments in female flowers or fully degenerated; filaments short, inserted between lobes of disk; anthers cordate or ovate. Ovary 4-carpellate; carpels free, ovoid, with 1 ovule; styles free or basally connate. Fruit comprising 1 or more drupelike monocarps (each a druparium), hard, somewhat fleshy. Seeds exalbuminous.
Shrubs or trees, monoecious, dioecious or bisexual. Leaves imparipinnate, exstipulate. Inflorescence cymose with axillary, bracteate, raceme-like thyrses. Sepals 4, fused at base. Petals 4, free, imbricate. Stamens 4, vestigial or absent in female flowers; filaments without basal appendage. Disc thick, 4-lobed. Carpels 4, free, vestigial in male flowers; styles free or cohering at base; ovule 1, pendulous. Fruit drupaceous; drupelets with thin pericarp and hard endocarp. Endosperm present.
Stamens (3) 4 (5), alternating with the petals, smaller in female flowers, inserted beneath and between the lobes of the disk, filaments free, unappendaged, glabrous.
Gynoecium of 4 free carpels, or carpels joined only at the base, absent or vestigial in male flowers; carpels 1-ovulate, ovule pendulous; styles (3) 4 (5), free.
Leaves sometimes ± crowded at the ends of the branches but often not crowded, usually imparipinnate; leaflets opposite, entire or toothed.
Inflorescences axillary, of spiciform panicles, andromonoecious (? or polygamous), male inflorescence larger than the female.
Flowers very small, usually in glomerules ± distantly arranged on the branches of the inflorescence.
Petals (3) 4 (5), longer than the calyx-lobes, imbricate.
Shrubs or trees, ± velvety-ferruginous-pubescent.
Calyx of (3) 4 (5) imbricate almost free lobes.
Fruit of 1–4 drupaceous mericarps.
Disk ± fleshy, (3) 4 (S)-lobed.
