Capparaceae Juss.

Family

Angiosperms > Brassicales

Characteristics

Shrubs or trees (deciduous or evergreen); spines absent (usually without thorns except Atamisquea); glabrous or puberulent with stellate trichomes or lepidote scales (producing glucosinolates). Stems erect or spreading; sparsely to profusely branched. Leaves alternate and distichous or spirally arranged, simple [palmately compound]; venation pinnate; stipules caducous, scalelike or absent; petioles present (pulvinus absent, nectaries present or absent, [petiolar spines present]); blade margins entire. Inflorescences terminal or axillary, usually racemose, sometimes corymbose or flowers solitary; pedunculate; bud scales usually persistent; bracts absent. Pedicels present. Flowers bisexual (sometimes appearing unisexual), actinomorphic or slightly zygomorphic, rotate to crateriform, campanulate, or urceolate; perianth and androecium hypogynous; sepals usually persistent (deciduous in Quadrella incana), 4, distinct; petals 4, attached directly to receptacle, imbricate, distinct, equal; intrastaminal nectary-discs or glands present or absent; stamens 6-250; filaments distinct, glabrous or pubescent; anthers dehiscing by longitudinal slits, pollen shed in single grains, binucleate, commonly tricolporate; pistil 1; ovary 1-carpellate, 2-locular; placentation parietal; ovules anatropous, bitegmic; style 1, (straight, relatively short and thick); stigma 1, capitate, unlobed. Fruits capsules or berries, valvate, elongate, dehiscent or not by 2 lateral valves (stipitate from elongation of gynophore). Seeds 1-38 or many, usually tan to yellowish brown or brown, sometimes green; arillate or not; endosperm scanty or absent, sometimes a persistent perisperm present; cotyledons incumbent to accumbent, (radicle-hypocotyl relatively short and conical). x = 8, 10.
More
Shrubs, trees, or woody vines, evergreen (deciduous in some Crateva), with branched or simple trichomes. Stipules spinelike, small, or absent. Leaves alternate or rarely opposite, simple or compound with 3[-9] leaflets. Inflorescences axillary or superaxillary, racemose, corymbose, subumbellate, or paniculate, 2-10-flowered or 1-flowered in leaf axil. Flowers bisexual or sometimes unisexual, actinomorphic or zygomorphic, often with caducous bracteoles. Sepals 4(-8), in 1 or 2 whorls, equal or not, distinct or basally connate, rarely outer whorl or all sepals connected and forming a cap. Petals (0-)4(-8), alternating with sepals, distinct, with or without a claw. Receptacle flat or tapered, often extended into an androgynophore, with nectar gland. Stamens (4-)6 to ca. 200; filaments on receptacle or androgynophore apex, distinct, inflexed or spiraled in bud; anthers basifixed (dorsifixed in Stixis), 2-celled, introrse, longitudinally dehiscent. Pistil 2(-8)-carpellate; gynophore ± as long as stamens; ovary ovoid and terete (linear and ridged in Borthwickia), 1-loculed, with 2 to several parietal placentae (3-6-loculed with axile placentation in Borthwickia and Stixis); ovules several to many, 2-tegmic; style obsolete or highly reduced, sometimes elongated and slender; stigma capitate or not obvious, rarely 3-branched. Fruit a berry or capsule, globose, ellipsoid, or linear, with tough indehiscent exocarp or valvately dehiscent. Seeds 1 to many per fruit, reniform to polygonal, smooth or with various sculpturing; embryo curved; endosperm small or absent.
Herbs, shrubs or trees, sometimes scandent, glabrous, pubescent, glandular or scabrous, occasionally lepidote. Leaves alternate or rarely opposite, simple or digitately 2–7-foliolate; leaflets entire, rarely serrate, dentate or panduriform. Stipules present, minute or spiny, caducous or persistent. Inflorescences terminal or axillary, racemose, often corymbose, or the flowers solitary or fascicled, often showy. Flowers hermaphrodite or very rarely dioecious, regular or irregular. Sepals 4–8, free or variously connate, valvate, imbricate or open in aestivation; receptacle cupular, funnel-shaped or cylindric, with an entire, undulate, dentate or fimbriate margin, sometimes very short. Petals 4–16 or absent, sessile or clawed. Stamens few to many, usually borne on a short or elongated androphore, all fertile or some without anthers; filaments free, equal or unequal, filiform; anthers 2-celled, oblong, longitudinally dehiscent. Ovary sessile or more usually supported on a gynophore of varying length, 1-locular with parietal placentas or divided by spurious dissepiments into 2 or more locules; ovules few to many. Fruit a capsule or berry, mostly oblong-cylindrical, globose or torulose, few-to many-seeded, rarely 1-seeded. Seeds usually reniform or angular; endosperm none or scanty; embryo arcuate or incurved; cotyledons folded or convolute
Herbs, shrubs, climbers or trees, glabrous or hairy; hairs sometimes glandular. Leaves alternate, petiolate, simple, or dissected or compound, sometimes heteroblastic; stipules spiny or absent. Inflorescence racemose, terminal or lateral; bracts small, caducous or absent. Flowers actinomorphic, sometimes slightly zygomorphic, bisexual, except Apophyllum. Sepals 4, rarely 2, in two whorls, equal or unequal, the outer whorl free or fused. Petals 4, rarely 2, equal, free. Stamens 1 to many, free, sometimes connate at base and adnate to gynophore; anthers bilocular, dehiscing by longitudinal slits. Receptacle sometimes with nectariferous glands. Ovary superior, 2–6-carpellate, 1–3-locular, sessile or usually on a gynophore; stigma simple, sessile; ovules 1 to many; placentation parietal. Fruit a capsule or berry. Seeds 1 to many, mostly coiled-reniform; embryo curved or coiled; testa smooth or sculptured, with or without aril.
Ovary usually borne on a more or less elongated gynophore, usually 1-locular with 2 parietal placentas but sometimes 2-locular by the intrusion of the placentas or multi-locular; ovules 4–?
Receptacle cupular, funnel-shaped or cylindric with an entire, undulate, dentate or fimbriate margin, sometimes very short
Inflorescences terminal or axillary, of racemes, corymbs or panicles, or flowers solitary and axillary
Flowers actinomorphic or zygomorphic, bisexual or unisexual by abortion, usually 4-merous, hypogynous
Leaves alternate, simple or digitately 3––9-foliolate; stipules absent or rudimentary, rarely spiny
Stamens 5–? usually borne on a short or elongated androphore, sometimes accompanied by staminodes
Seeds reniform or subglobose, without endosperm; embryo usually curved
Herbs, shrubs, trees or lianes
Fruit a capsule or a berry
Petals 4 (0, 5, 6 or more)
Style short or absent
Sepals 3–4 (5)
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Foliage retention evergreen
Sexuality hermaphrodite
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Images

Capparaceae unspecified picture
Capparaceae unspecified picture

Distribution

Capparaceae world distribution map, present in Australia and China

Identifiers

LSID urn:lsid:ipni.org:names:30001562-2
WFO ID wfo-7000000111
COL ID 7PH
BDTFX ID 101011
INPN ID 187297
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Capparaceae

Lower taxons

Boscia Capparis Neothorelia Poilanedora Buchholzia Crateva Morisonia Roydsia Beautempsia Bachmannia Belencita Cadaba Maerua Ritchiea Acome Thilachium Hypselandra Neocalyptrocalyx