Crateva L.

Crateva (en)

Genus

Angiosperms > Brassicales > Capparaceae

Characteristics

Small to medium-sized trees, facultatively shortly deciduous and then flowering when bare; glabrous. Branchlets terete with distinct leaf-scars. Stipules small, caducous. Leaves 3-foliolate, the top of the long petiole sometimes bearing glandlike appendages on the upper surface. Leaflets sessile to shortly stalked, the lateral ones basiscopically asymmetrical, sometimes with more or less distinct pellucid dots. Raceme terminal, corymbiform, either with arrested growth or growing through and developing into a leafy twig with lateral flowers. Flowers sustained by bracts, rarely by leaves, pedicelled, opening at a very early stage of development, floral parts not persistent. Bracts stipulate. Receptacle wide; disk dish-shaped and incurved. Sepals equal, ovate-spathulate, green. Petals equal, unguiculate, more or less ovate to rhomboid with narrowed base, first white, later cream-coloured, the lower (anterior) pair tending to take a transversal (horizontal) position. Stamens (8-)12-30, filaments at the very base connate with the gynophore, long, filiform, spreading. Gynophore approximately as long as the stamens. Ovary 1-locular, the 2 placentas sometimes intruding to about halfway the lumen but not coalescent. Stigma conspicuous, flat, soon after anthesis obsolete. In fruit pedicel, torus and gynophore woody and more or less thickened, the last with a whorl of filament-scars near the base; the gynophore mostly not stretching. Berry large, 1-celled, with tough, sometimes papillate skin. Seeds densely packed; embedded in pulp, horseshoe-shaped, smooth or crested, one cotyledon larger, curved round the other.
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Trees or sometimes shrubs, evergreen or deciduous, glabrous throughout. Twigs terete or angular, with lenticels, with pith or hollow. Stipules triangular, small, caducous. Leaves alternate, palmately compound; petiole long, distal portion near rachis often with glands; leaflets 3, disarticulating from petiole; petiolules short, thin when young, becoming thick in maturity; lateral leaflet blades with asymmetric base. Inflorescences at tip of new branches, corymbose racemes; rachis either with arrested growth after anthesis or growing through a leafy twig, often with obvious scars after pedicels fall; bract at base of pedicels, caducous. Flowers bisexual or unisexual by failure of one sex to develop. Pedicel long. Receptacle disklike, inner surface concave, with nectary, with sepal and petals on margin. Sepals 4, greenish, equal, obviously smaller than petals, deciduous. Petals 4, white, cream-colored, or yellow, equal, clawed, blade ovate to rhomboid with 4-6 secondary veins on each side of midvein. Stamens (8-)12-50; filaments basally connate to form a 1-4 mm androgynophore. Gynophore 2-8 cm but degenerated in staminate flower; ovary 1-locular, placentae 2, ovules many; style short or absent; stigma inconspicuous, knob-shaped. Fruit a berry, globose or ellipsoid, drooping; pericarp drying to gray, red, purple, or brown, leathery, firm, apically smooth or papillate; fruiting pedicel, receptacle, and gynophore woody and thickened. Seeds 25-50 per berry, embedded in creamy fetid or pungent mesocarp; seed coat smooth; cotyledons convolute, one longer and curved around other; radicle conical, short.
Small trees with pedately 3-foliolate, membranous leaves. Inflorescence a terminal, pseudo-terminal or lateral often corymbose raceme, apparently at least occasionally growing out as a leafy twig; bracts 3-foliolate or subulate. Sepals 4, irregularly imbricate, the buds opening prematurely to expose the immature corolla, stamens and gynoecium; receptacle shallowand bowl-shaped, the thickened or inflexed rim forming the disk. Petals 4, clawed. Androphore short and stout, scarcely as long as the receptacle; stamens 15–20. Ovary globose with 2 lateral placentas, many-ovulate. Fruits globose, crustaceous, many-seeded.
Trees, glabrous. Leaves compound, trifoliolate; petiole often glandular; stipules minute, triangular, caducous. Inflorescence corymbose; flowers bracteate. Sepals equal, valvate. Petals subequal, long-clawed. Stamens 8–50; filaments arising from basal androgynophore. Receptacle dish-shaped, nectariferous. Gynophore slender; ovary 1-locular; placentas 2. Fruit a globose, leathery berry. Seeds large, horseshoe-shaped.
Ovary ovoid or globose on a long gynophore, 1-locular or 2-locular by intrusion of the 2 placentas, multi-ovulate; stigma subsessile.
Fruit globose or ovoid with a coriaceous pericarp, borne on a stout stipe.
Sepals 4, open in aestivation, arising from a shallow receptacle.
Inflorescence of axillary or terminal, corymbose racemes.
Shrubs or trees with 3-foliolate, petiolate leaves.
Seeds many, reniform with a coriaceous testa.
Flowers bisexual or unisexual by abortion.
Petals 4 (5), rather large, clawed.
Androgynophore short, dilated.
Stamens 8–50.
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Growth form tree
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Foliage retention evergreen
Sexuality hermaphrodite
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Environment

Mostly in periodically inundated lowland forest near rivers, below 700 m. In dry regions shortly deciduous, the flowers then appearing simultaneously with the flush. Also cultivated for ornamental purposes and presumably occasionally introduced.
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Hardiness (USDA) 8-12

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Cultivation

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