Caryota L.

Fishtail palm (en)

Genus

Angiosperms > Arecales > Arecaceae

Characteristics

Stems small to very large, solitary or clustered, columnar or swollen, usually ringed with conspicuous leaf scars. Leaves 4-20, spread out along stem, or borne in a compact crown at top of stem; leaf sheaths closed, not forming crownshafts; petioles short or elongate and then rounded in cross section, covered with whitish or brownish hairs, sometimes striped; leaves bipinnate, each primary pinna made up of several secondary pinnae borne on a secondary rachis terminated by a pinna; individual secondary pinnae triangular, with jagged outer margins, often spreading in different planes. Plants monoecious, semelparous, flowering proceeding from top of stem downward (basipetal). Inflorescences usually branched to 1 order, rarely spicate, borne either among or below leaves, covered with many persistent bracts; peduncle bearing a prophyll and several peduncular bracts; rachillae usually many, long and pendulous; flowers unisexual, borne in threes of 1 central female and 2 lateral male; male petals either yellow or purplish; stamens 6-150. Fruits orange, red, or purple, medium sized, usually ± globose, 1-or 2-seeded; mesocarp filled with irritant needle-shaped crystals; endosperm ruminate (homogeneous in one species); germination remote; eophylls bifid [pinnate in one species] with jagged margins.
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Stems solitary or clustered, slender to massive, smooth, with conspicuous nodal rings. Leaves: blade 2-pinnate (1-pinnate in juvenile plants); plication induplicate; segments cuneate, in 1 plane; apices jagged and irregular; basal segments not modified into spines. Inflorescences initiated basipetally, first one appearing terminal, successive one borne axillary among leaves, and later ones below leaves, pendulous, paniculate, with 1 order of branching [spicate]; prophyll small; peduncular bracts numerous, tubular. Flowers unisexual, sessile, borne in triads of 1 pistillate flower flanked by 2 staminate flowers. Staminate flowers: sepals 3, imbricate, free; petals 3, connate basally, valvate; stamens numerous [6], free; pistillode absent. Pistillate flowers: sepals 3, imbricate, free; petals 3, connate for nearly 1/2 length, valvate; staminodial lobes present or absent; pistils 1, 3-loculate; ovules 1 per locule; styles indistinct; stigmas 3-lobed. Fruits berries, globular; exocarp purple, smooth; mesocarp fleshy, containing irritating raphides; endocarp absent. Seeds globular; endosperm ruminate [homogeneous]; embryo lateral; eophyll 2-cleft, segments fan-shaped. n = 17.
Clustering (not in Australia) or solitary, monoecious, unarmed, hapaxanthic palms. Leaves bipinnate. Pinnae induplicate, cuneate, apically praemorse, truncate. Inflorescence spicate (not in Australia) or branched to 1 or more orders, produced basipetally from axillary nodes; prophyll inconspicuous; peduncular bracts several, spirally arranged on short peduncle. Flowers in triads, most frequently proximal on rachilla, with single or paired staminate flowers in distal portion. Staminate flower with imbricate sepals; petals valvate; stamens few–many; pistillode absent. Pistillate flower with petals basally connate; staminodes sometimes present; ovary trilocular with 1–3 fertile locules. Fruit globose (not in Australia) to sub-globose, 1–3-seeded; stigmatic remains apical, or lateral (not in Australia); mesocarp with irritant crystals; endocarp not differentiated. Seed tear-shaped (not in Australia), hemispherical or globose, smooth, black; endosperm ruminate or homogeneous (not in Australia); embryo lateral (not in Australia) or sub-basal. Eophyll bifid.
Life form perennial
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Foliage retention deciduous
Sexuality monoecy
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Hardiness (USDA) 8-12

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