Gleicheniaceae C.Presl

Family

Pteridophytes > Gleicheniales

Characteristics

Plants coarse, terrestrial. Stems long-creeping, forked, stele protostelic (a solid rod of vascular tissue with phloem surrounding xylem), covered with scales or hairs. Leaves monomorphic, large, scrambling or trailing, 1--many times forked. Petiole not articulate to stem, with pair of opposite pinnae and arrested bud at apex, or rachis continuing and producing 2 or more pairs of opposite pinnae. Pinnae 1--several times forked, with arrested bud at each fork; indument of simple, branched, or stellate hairs [scales]. Veins free, 1--4-forked. Sori round, indusia absent. Sporangia 2--many on slightly elevated receptacle, sessile to subsessile; annulus complete, transverse, medial, longitudinally dehiscent; spores 120--800 per sporangium. Spores all alike, whitish to yellowish, bilateral to globose, monolete or trilete, generally smooth without elaborately ornamented surface. Gametophyte borne aboveground, green, obcordate to elongate.
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Plants terrestrial, often forming thickets. Rhizomes long creeping, protostelic, with scales or multicellular hairs. Fronds monomorphic, evergreen, erect, scrambling, or climbing, vernation circinate; stipe not articulate, distant, cylindrical, apex forked, with dormant bud; apical bud covered with hairs or scales; rachis simple to several times pseudodichotomously branched [or pinnate], with stellate hairs and/or ciliate scales when young, these persistent or glabrescent; pinnules pectinately pinnatisect; lobes elliptic or lanceolate, apex obtuse or acute; veins free, forked; lamina papery or subleathery, frequently glaucous abaxially. Sori in 1(-3) lines on both sides of lobe costules, orbicular, exindusiate; sporangia 2-10[-16], sessile, turbinate, with a mid-transverse annulus; spores tetrahedral or reniform, without perispore, smooth.
Terrestrial homosporous ferns. Rhizome slender, protostelic, rarely solenostelic in Dicranopteris (not in Australia), bearing scales or hairs. Fronds erect, scrambling or climbing; lamina pseudodichotomously branched, once or more pinnate, of indeterminate growth due to presence of dormant buds in branch axils (absent in Gleichenia abscida ); segments adnate; veins free; vernation circinnate. Sori round or nearly so, exindusiate, consisting of a small number of large sporangia with an oblique or nearly apical annulus. Spores without chlorophyll. Gametophyte long-lived, elongate to subcordate.
Perennial terrestrial herbs with wide–creeping rhizomes; frond apparently dichotomous, due to arrested growth owing to the formation of a dormant terminal bud; apex of rhizome and buds covered with scales or hairs; stipule–like leaflets present in some species; pinnae lobed almost to the costa, elongate (in W.African species), with forked free veins; sori on the veins punctate with a few large subsessile sporangia; no indusium; annulus uniseriate, sharply defined, oblique and complete except at the line of dehiscence; dehiscence vertical; spore trilete (in W.African species)
Terrestrial herbs, often large, with widely creeping rhizomes bearing scales and/or hairs
Life form perennial
Growth form herb
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Foliage retention evergreen
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Root system rhizome
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Environment

The family as a whole seems to favour severely disturbed or pioneer habitats, often at rainforest margins, and frequently in mineral-poor substrates. The taxa are not widely nor easily cultivated, probably due to their specific habitat requirements.
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Hardiness (USDA) 7-11

Usage

Uses -
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Cultivation

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Images

Gleicheniaceae unspecified picture

Distribution

Gleicheniaceae world distribution map, present in Australia and China

Identifiers

LSID urn:lsid:ipni.org:names:30003211-2
WFO ID wfo-7000000251
COL ID -
BDTFX ID -
INPN ID 445327
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Gleicheniaceae Stromatopteridaceae

Lower taxons

Gleichenella Sticherus Stromatopteris Diplopterygium Dicranopteris Gleichenia