Shrubs [trees], monoecious [dioecious], hairy [glabrous], hairs simple; branching phyllanthoid. Leaves deciduous [persistent], alternate, simple, scalelike on main stems, well developed on ultimate branchlets; stipules persistent; blade margins entire. Inflorescences unisexual or bisexual, fascicles [cymules], proximal mostly or entirely staminate, distal mostly or entirely pistillate. Pedicels present [absent]. Staminate flowers: sepals [5–]6, distinct; petals 0; nectary absent; stamens 3–4[–8]; filaments and anthers connate; connectives extending beyond anthers as deltoid appendages; pistillode absent. Pistillate flowers: sepals persistent, 6, distinct; petals 0; nectary absent; pistil [3–]5–10[–15]-carpellate; styles [3–]5–10[–15], connate nearly to apex, unbranched. Fruits capsules. Seeds 2 per locule, rounded-trigonous [ellipsoid]; seed coat ± fleshy, smooth; caruncle absent. x = 13, 16.