Gouania Jacq.

Chewstick (en)

Genus

Angiosperms > Rosales > Rhamnaceae

Characteristics

Climbing shrubs or lianes with circinnate tendrils (rarely, or apparently never in Africa, seeming to be mere shrubs). Leaves alternate, petiolate; blades usually ovate, serrate (rarely nearly entire), acute or acuminate, at base usually rounded or cordate and often 3–5-nerved but penninerved more distally. Stipules present, free, usually caducous. Cymes usually small, but usually aggregated into ample leafless terminal panicles. Flowers 5-merous, bisexual, epigynous at all stages. Disk rather massive, epigynous. Ovary usually urceolate; style trifid or obscurely 3-lobed. Fruit a dry schizocarpous capsule, late in maturation the endo-and exocarpous tissue near the septa (on both sides of each mericarp) producing 3 broad thin wings (thus each wing is composed of parts of 2 adjacent mericarps; reportedly the wing-production fails in certain Asian and American, but not African, populations, and perhaps the reports are based on observations of immature fruit, for the wing-proliferation is a quite late development).
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Shrubs climbing, usually with tendrils, unarmed. Leaves alternate, with caducous stipules, petiolate, pinnativeined or 3-veined from base, margin entire or dentate. Flowers polygamous, in terminal or axillary, cymose racemes or cymose panicles, often with tendrils at lower part of rachis or base. Calyx tube short, coadnate with ovary; sepals 5, ovate-triangular, adaxially medially keeled, dehiscent with longitudinal slits. Petals 5, spatulate. Stamens 5, dorsifixed, enfolded by petals. Disk thick, pentagonous or 5-lobed, glabrous or hairy. Ovary inferior, ± deeply immersed in disk, 3-loculed, with 1 ovule per locule; styles cleft to half or deeply cleft. Capsule subglobose, impressed on both ends, crowned by persistent limb of calyx, 3-winged, with 3 rounded locules split from axil at maturity; locules indehiscent or along inner rib narrowly fissured. Seeds 3, red-brown, shiny, obovoid; endosperm thin.
Lianas, or scandent to arching shrubs, climbing by tendrils. Leaves alternate, pinnately-nerved or sometimes 3-nerved from the base, mostly dentate, the teeth frequently glandular; petiolate, the stipules narrow and deciduous or sometimes broad and persistent. Inflorescences spicate thyrses composed of glomerules, axillary or terminal. Flowers bisexual or 'polygamous; floral tube obconic to siibcampanulate; sepals 5, persistent; petals 5, white to green white, short-clawed; stamens 5, hidden by the petals; nectariferous disc 5-lobed or pentagonal, the lobes opposite the sepals; ovary inferior, immersed in the disc, 3-locular, the style 3-branched more or less basally, the stigmas minute. Fruit a schizocarp, 3-locular, generally 3-winged, splitting longitudinally along the margin of each wing into 3, 2-winged mericarps; seeds 3, the seed coat shiny.
Lianes, tall, climbing by curled shoot-tips, or (not in Australia) shrubs, simple-pubescent. Leaves alternate, petiolate, discolorous or ± concolourous, penniveined; stipules free, caducous. Inflorescences axillary or terminal, elongate pseudoracemes (these sometimes arranged in panicles) with flowers in contracted cymose clusters on the axes; bracts persistent or caducous. Flowers bisexual, 5-merous, greenish. Hypanthium cup-shaped. Sepals incurved to erect, persistent on the fruits. Petals cucullate, obscurely clawed, incurved. Stamens subequal to the petals, incurved. Disc conspicuous, lining the hypanthium, often lobed, smooth, glabrous. Ovary immersed in disc; carpels 3 (or 4); style branched. Fruit a dry, laterally-winged schizocarp, splitting septicidally into 3 (or 4) two-winged, one-seeded articles; torus apical.
Woody vines [climbing shrubs], tendrils present [rarely absent], unarmed; bud scales present. Leaves deciduous [persistent], alternate; blade not gland-dotted; pinnately veined [3-veined from base]. Inflorescences: axillary or terminal, racemelike or paniclelike thyrses; peduncles and pedicels not fleshy in fruit. Pedicels present or absent. Flowers bisexual [unisexual, plants polygamous]; hypanthium cupulate to campanulate, 1.5–3 mm wide; sepals 5, spreading, pale green, ovate-triangular, keeled adaxially; petals 5 [rarely 0], white to yellowish or greenish, hooded, spatulate, clawed; nectary fleshy, margins 5-lobed [unlobed], lining hypanthium but distally free; stamens 5, enfolded by petals; ovary inferior, 3-locular; styles 3, connate basally. Fruits schizocarps, breaking into 3 2-winged samaras [rarely nutlets].
Fruit a schizocarp, longitudinally 3-winged (or 3-angled), septicidal, separating into 3 woody or coriaceous indehiscent cocci.
Leaves alternate, petiolate; lamina entire or dentate, penninerved but often 3–5-nerved from the base.
Ovary inferior, immersed in the disk; style 3-fid or obscurely 3-lobed.
Climbing shrubs or lianes; branches provided with coiled tendrils.
Petals 5, cucullate; inserted below the margin of the disk.
Disk filling the receptacle, 5-lobed.
Inflorescence a paniculate thyrse.
Seeds compressed or planoconvex.
Stipules small, caducous.
Flowers usually bisexual.
Stamens 5.
Sepals 5.
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Growth form shrub
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Foliage retention deciduous
Sexuality hermaphrodite
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Environment

Rainforest margins from near sea level to c. 250 m alt. (rarely higher).
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Hardiness (USDA) 8-12

Usage

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Cultivation

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Images

Gouania unspecified picture

Distribution

Gouania world distribution map, present in Australia, China, Panama, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:30010171-2
WFO ID wfo-4000016086
COL ID 4QGM
BDTFX ID -
INPN ID 445913
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Pleuranthodes Gouania

Lower taxons

Gouania aphrodes Gouania cornifolia Gouania ekmanii Gouania eurycarpa Gouania hillebrandii Gouania javanica Gouania latifolia Gouania laxiflora Gouania leratii Gouania lineata Gouania longipetala Gouania longispicata Gouania lupuloides Gouania mangarevica Gouania mauritiana Gouania meyenii Gouania polygama Gouania rosei Gouania stipularis Gouania tiliifolia Gouania ulugurica Gouania virgata Gouania viridis Gouania vitifolia Gouania wurdackii Gouania acalyphoides Gouania acreana Gouania aptera Gouania blanchetiana Gouania canescens Gouania colombiana Gouania conzattii Gouania corylifolia Gouania frangulifolia Gouania humbertii Gouania inornata Gouania microcarpa Gouania myriocarpa Gouania pannigera Gouania pyrifolia Gouania richii Gouania riparia Gouania rumicina Gouania striata Gouania trichodonta Gouania ulmifolia Gouania phillipsonii Gouania zebrifolia Gouania taolagnarensis Gouania perrieri Gouania callmanderi Gouania gautieri Gouania cupreifolia Gouania ambrensis Gouania cupuliflora Gouania croatii Gouania ferruginea Gouania guiengolensis Gouania obamana Gouania pubidisca Gouania andamanica Gouania australiana Gouania efatensis Gouania exilis Gouania hillii Gouania leptostachya Gouania alnifolia Gouania discolor Gouania napalensis Gouania velutina Gouania hypoglauca