Trees or shrubs, the Panamanian species all generally tree-like, but may be shrubby from stump sprouts. Leaves pinnate-compound, only rarely with a terminal leaflet, the leaflets opposite or alternate. Inflorescences axillary. Flowers perfect; calyx saucer to cup-shaped or of 5 nearly distinct sepals; petals 4-6; staminal tube urceolate or cylindric, entire or shallowly lobed, anthers 8-12, included or, rarely, barely exserted;-ovary 4-to 12-celled, the ovules 1 or 2 per cell, superposed. Fruit a thick or woody-valved capsule loculicidally dehiscent (sometimes tardily so) from the apex; seeds 1 per capsule or 1-2 per locule, more or less arillate, cotyledons fleshy.