Lianas, rarely shrubs or climbing shrubs. Leaves opposite, rarely ternate (not Panama), simple, entire, petiolate or subsessile, coriaceous, chartaceous, or papyraceous; stipules conspicuous, adnate to the petioles. Inflorescences of sim-ple, many-flowered umbels, or with the umbels arranged in cymose panicles; pedicels sessile, conspicuously bracteolate at the base. Flowers with the sepals ovate or ovate-oblong, 8-10, glandular or eglandular; petals clawed, yellow or white, rarely reddish (not Panama), generally glabrous, the limb suborbicular, obovate or ovate, concave, subentire, denticulate or shortly fimbriate, the fifth petal smaller, smooth, fimbriate or glandulose; stamens 10, somewhat unequal, glabrous, the filaments united at the base, the anthers ellipsoid or sometimes oblong; style glabrous with the apex truncate and the stigma lateral, erect, the apex scarcely thickened or more or less sigmoid with the apex dilated and the posterior angle apiculate, the ovary trilobate, hirsute. Fruit a schizocarp of 3 samaras, each with 2 lateral wings and a short dorsal ridge or crest, sometimes also with short appendages between the lateral wings and the dorsal crest, the lateral wings usually rounded or flabellate; cotyledons unequal, the outer 1 larger and enveloping the inner 1.