Phyla Lour.

Fogfruit (en), Verveine (fr)

Genus

Angiosperms > Lamiales > Verbenaceae

Characteristics

Perennial mostly procumbent or creeping herbs, stems trailing or ascending, sometimes slightly woody basally, more or less incanous throughout with ap-pressed-strigose, rarely subglabrous malpighian (medifixed) trichomes. Leaves decussate-opposite, often fleshy, flat or pinnately plicatulate. Inflorescences spicate, axillary, cylindric, densely many-flowered, usually greatly elongate in fruit, solitary, paired, or ternate, never aggregated into corymbs or panicles. Flotvers small, sessile, borne singly in the axils of small cuneate-obovate or flabel-liform bractlets, not at all 4-ranked; calyx small, membranous, ovoid-campanulate or compressed and 2-carinate or winged, the rim 2-or 4-fid or 4-dentate; corolla irregular, the tube straight or incurved, slender, slightly exserted from the calyx, equal in diameter throughout or slightly ampliate above, the limb oblique, spread-ing, somewhat 2-lipped, 4-parted, the lobes broad, often retuse apically, the posterior entire, emarginate, or bifid to about the middle, the laterals exterior, the anterior often larger; stamens 4, didynamous, included or slightly exserted, the anthers unappendaged; ovary 2-loculed, the ovules 1 per locule, the stigma incrassate, oblique or recurved. Fruit small, dry, included by the calyx and sometimes adnate to it, dividing into 2 pyrenes at maturity, the pericarp leathery and hard, the exocarp membranous and rarely distinct from the pyrenes.
More
Perennial herbs, procumbent or creeping; tomentum of medifixed hairs. Stem trailing or ascending, often rooting at nodes. Leaves simple, decussate, petiolate or subsessile. Inflorescence spicate, axillary, pedunculate. Flowers bracteate, sessile, zygomorphic; bracts imbricate. Calyx persistent, membranous, ovoid to campanulate or compressed, with 2 keels or wings and 2-lobed; rim 2-or 4-fid or 4-dentate. Corolla hypocrateriform; tube cylindrical or slightly dilated upwards; lobes obliquely subbilabiate, 4–5-fid, upper lip entire to 2-lobed, lower lip 3-fid. Stamens 4, didynamous, epipetalous, included or scarcely exserted; anthers nonappendiculate. Ovary 2-locular, with 1 ovule in each locule; style short; stigma oblique, thick. Fruit schizocarpic, splitting into two 1-seeded mericarps. Seeds without endosperm.
Fls in dense, elongating, pedunculate axillary spikes; bracts small, cuneate-obovate to flabelliform; cal small, membranous, often compressed or even winged, with 2 or 4 teeth or segments; cor-tube very slender, straight or incurved, slightly exserted from the cal, the limb oblique, spreading, 4-parted and somewhat 2-lipped; stamens 4, paired, included or slightly exserted; anthers unappendaged; ovary with 2 uniovulate chambers; stigma thickened, oblique or recurved; fr included in the cal or even adnate to it, dividing into 2 nutlets, sometimes with separating thin exocarp; prostrate or creeping perennials with ± ascending or trailing branches, ± canescent with malpighian hairs. 10, originally N. Amer.
Trailing perennial herbs with angular stems rooting at the nodes. Leaves opposite, simple, subentire to toothed, mostly tapering into an obscure petiole; lamina with sparse to dense adpressed medifixed hairs. Inflorescences ± long-pedunculate axillary short ovoid to cylindric very dense bracteate spikes. Calyx membranous, 2-lobed, compressed. Corolla with short tube and spreading somewhat irregularly 4-lobed white or purple limb. Stamens 4, included. Fruit of 2 nutlets enveloped by the persistent calyx.
Herbs. Branches usually acutely 4-angled, sometimes spiny, hirsute, glandular. Leaves opposite. Inflorescences terminal, spicate or capitate, elongated in fruit; bracts obovate. Flowers small. Calyx membranous, 2-lipped, 4-lobed. Corolla sometimes 2-lipped; lobes 5, spreading. Stamens 4, didynamous, inserted on distal part of corolla tube, included. Ovary 2-locular; ovule 1 per locule. Style short; stigma capitate. Capsules small, splitting into 2 1-seeded cocci.
Mat-forming, unarmed herbs with prostrate, creeping stems. Lvs opposite, toothed or lobed, with medifixed hairs. Infl. a dense, subglobose to cylindric, axillary spike; fls sessile, small, bracteate. Calyx 2-lobed, divided almost to base, ± accrescent. Corolla limb 4-lobed, with 2 unequal lips; tube > calyx, straight. Stamens 4, included, didynamous. Ovary 2-celled, each cell with 1 ovule. Style included. Fr. dry, separating into 2 nutlets.
Corolla usually white, becoming rose-coloured; tube straight or curved, cylindric or widening toward the apex; limb oblique, c. 2-lipped, with the (upper) adaxial lip entire, emarginate or 2-lobed, and the lower (abaxial) lip 3-lobed with the median lobe usually larger than the others.
Fruit a schizocarp, compressed oblong, separating at maturity into two 1-locular mericarps; mericarps with a dry hard pericarp, semi ovoid, flat on the commissural face, rounded and smooth on the dorsal face, glabrous.
Inflorescences axillary, pedunculate, spicate; spikes short, dense, ovoid becoming cylindric after anthesis, with bracts closely imbricate.
Prostrate perennial herbs with woody taproots; indumentum of sparse to closely spaced appressed medifixed hairs or plants glabrous.
Ovary 1-carpellate, carpel 2-locular, each locule 1-ovulate; style short, undivided; stigma thick, obliquely capitate.
Leaves opposite, simple, shortly petiolate or subsessile, serrate.
Stems trailing, rooting at the nodes; branches short, ascending.
Calyx membranous, dorsiventrally flattened, 2-lobed, persistent.
Flowers small, sessile, each solitary in the axil of a bract.
Stamens 4, included or somewhat exserted.
Life form perennial
Growth form herb
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Sexuality hermaphrodite
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Root system tap-root
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Environment

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Hardiness (USDA) 6-12

Usage

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Edible -
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Cultivation

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Images

Phyla unspecified picture

Distribution

Phyla world distribution map, present in Aruba, Angola, Åland Islands, Albania, Andorra, Australia, Benin, Burkina Faso, Bangladesh, Bahamas, Belize, Bermuda, Bolivia (Plurinational State of), Botswana, Central African Republic, China, Colombia, Cabo Verde, Costa Rica, Cuba, Cayman Islands, Cyprus, Djibouti, Dominica, Algeria, Ecuador, Egypt, Eritrea, Spain, Ethiopia, France, Micronesia (Federated States of), Georgia, Ghana, Guinea-Bissau, Greece, Guatemala, Guyana, Honduras, Haiti, Indonesia, Iran (Islamic Republic of), Italy, Jamaica, Japan, Kenya, Liberia, Libya, Lesotho, Morocco, Madagascar, Maldives, Mali, Myanmar, Mozambique, Mauritania, Mauritius, Namibia, Nigeria, Nicaragua, Nepal, New Zealand, Oman, Pakistan, Panama, Peru, Philippines, Palau, Puerto Rico, Portugal, Paraguay, Saudi Arabia, Sudan, Senegal, Singapore, Solomon Islands, Somalia, Suriname, eSwatini, Seychelles, Chad, Togo, Thailand, Tunisia, Tanzania, United Republic of, Uganda, Uruguay, United States of America, Venezuela (Bolivarian Republic of), Samoa, Zambia, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:330571-2
WFO ID wfo-4000029368
COL ID 8VS9C
BDTFX ID 86985
INPN ID 196188
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Phyla

Lower taxons

Phyla betulifolia Phyla cuneifolia Phyla linearis Phyla x intermedia Phyla nodiflora Phyla lanceolata