Pyrola L.

Wintergreen (en), Pirole (fr), Pyrole (fr)

Genus

Angiosperms > Ericales > Ericaceae

Characteristics

Herbs, chlorophyllous, autotrophic (achlorophyllous and heterotrophic in forms of P. chlorantha and P. picta). Stems erect, glabrous. Leaves essentially basal or, sometimes, highly reduced or absent (P. chlorantha, P. picta), alternate; petiole present; blade maculate or not, elliptic, ovate-elliptic, oblong-elliptic, oblanceolate, oblong-obovate, ovate, obovate, spatulate, subreniform, reniform, or round, subcoriaceous to coriaceous, margins entire, denticulate, crenulate, crenate, or crenate-serrulate, plane or revolute, surfaces glabrous. Inflorescences racemes, usually erect in flower and fruit, (symmetric); peduncular bracts present or absent; inflorescence bracts free from pedicels. Pedicels pendent in fruit; bracteoles absent. Flowers radially symmetric (bilaterally symmetric in P. minor), spreading or nodding; sepals 5, connate proximally, often obscurely so, calyx lobes lanceolate, ovate, triangular, deltate, oblong, or obovate; petals 5, distinct, white, greenish white, yellowish white, pink, or purplish red, without basal tubercles, corolla crateriform to broadly campanulate; intrastaminal nectary disc absent; stamens 10, exserted; filaments broad proximally, gradually narrowed medially, slender distally, glabrous; anthers oblong, without awns, with or without tubules, dehiscent by 2 round to elliptic or obovate pores; pistil 5-carpellate; ovary imperfectly 5-locular; placentation intruded-parietal; style (exserted or included), bent downward or straight (P. minor), expanded distally; stigma 5-lobed, without subtending ring of hairs. Fruits capsular, pendulous, dehiscence loculicidal, cobwebby tissue exposed by splitting valves at dehiscence. Seeds ca. 1000, fusiform, winged. x = 23.
More
Perennial glabrous herbs or small half-shrubs, with rhizomes, short leafy stems. Leaves spiral, persistent, entire or denticulate, long-petioled. Flowers in terminal, simple, bracteate racemes, nodding, white (Mal.), actinomorphic or slightly zy-gomorphic. Calyx deeply 5-lobed, persistent. Petals 5, concave, sessile, ± arranged in form of a bell or somewhat spreading, caducous. Stamens 10; filaments subulate; anthers incompletely 4-celled, dehiscing by terminal pores. Ovary 5-celled; ovules ∞; style columnar, thickened upwards, straight (Mal.); stigma truncate or with a shortly 5-lobed stigma. Capsule depressed-subglobose, somewhat pentagonal, 5-celled, loculicidally 5-valved from the base, crowned by a persistent style. Seeds minute; testa lax, produced on both ends.
Herbs suffruticose, evergreen, erect, glabrous. Rootstock long, slender, branched, with sparse fine roots. Aerial stem ascending, not branched, scaly at base. Leaves in a rosette at base of caudex, long petiolate; leaf blade abaxially light green, adaxially green, margin entire or serrate. Scapes long, slender, with scalelike bracts at base. Flowers many, nodding, in racemes. Calyx 5-lobed, persistent. Stamens glabrous; anthers opening by pores. Ovary depressed-globose; style exserted, usually curved distally and dilated apically into a ring; stigma capitate, shortly 5-lobed. Capsules nodding, valves connected by fibers at valve margins. Seeds many, small, with thin testa prolonged at both ends. n = 23, 46.
Pet 5; filaments elongate, slender; anthers subapically attached, opening by terminal pores, the pollen sacs often prolonged above into short tubes; style ± elongate; capsule opening from the base upward; rhizomatous herbs, the few broad, petiolate lvs nearly basal, persistent throughout the winter (rarely the lvs suppressed); fls in an erect, terminal, long-peduncled raceme. 40/N. Temp.
Life form perennial
Growth form herb
Growth support -
Foliage retention evergreen
Sexuality hermaphrodite
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Environment

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Hardiness (USDA) 4-9

Usage

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Edible -
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Cultivation

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Germination duration (days) 30 - 90
Germination temperacture (C°) 12
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