Flowers dioecious, solitary or fasciculate in the ocreolae, short-pedicellate to subsessile, the pedicels, rhachises and dorsal surfaces of the ocreolae usually densely pubescent. Staminate flowers in fascicles of 1-5 subsessile pairs; tepals 6, sub-equal, linear to ovate, uniseriate, pubescent; stamens 9, exceeding the more or less campanulate tepals, the filaments discretely attached to the bases of the free portions of the tepals, not forming a distinct annular ring; anthers versatile, introrse, 4-locular; pistillodes absent. Pistillate flowers pedicellate and mostly solitary in the ochreolae; tepals in 2 strongly dissimilar series; outer 3 (sepals) connate at the bases forming a rhomboidal to campanulate tube, the distal portions developing into oblanceolate wings; inner 3 (petals) much smaller, linear to ovate, free or partially adnate to the tube; staminodia absent or occasionally forming an annular disc from which apparently functional stamens arise; ovary trigonous; styles 3, with verrucose stigmatic tissue extending down their inner surfaces to about half their length. Achenes included, triquetrous or rarely terete, with a small beak formed of the persistent style bases, brown, lustrous or dull, occasionally verrucose or punctate. Trees with glabrous to densely tomentose or strigose hollow branches. Leaves alternate, entire, glabrous to densely tomentose or strigose, oblong to ovate, often slightly inequilateral, often with conspicuous longitudinal striae representing the plications of the leaf in the bud, the veins immersed above, but prominent below; ochreae deciduous. Inflorescences of predominantly terminal subspicate panicles or racemes.
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