Dioecious shrubs or trees, the younger portions usually pubescent with stellate hairs, the sap often reddish or brownish. Leaves alternate, glabrous above, stellate-pubescent and often glabrescent below, submembranaceous to coriaceous, petiolate, exstipulate, the tertiary veins obscure. Flowers in fascicles, racemes or panicles, the axes usually stellate-pubescent; bracts membranaceous, deciduous; bracteoles absent. Staminate flowers in fascicles of 3-15, the perianth deeply or shallowly 3-to 4-parted; anthers 2-6, dorsally connate, at least basally, longer or shorter than the column. Pistillate flowers solitary or in fascicles of 2-7, the tepals partially connate. Fruits 1-40 per inflorescence, the ligneous pericarp dehiscing longitudinally into 2 valves; seed globose to ellipsoid, the aril laciniate.