Epiphytic or terrestrial, suffrutescent herbs or soft woody shrubs. Leaves sim-ple, exstipulate, alternate, petiolate, pinnately veined and usually denticulate or serrulate. Flowers solitary in the axils of the upper leaves on ebracteolate pedicels or more rarely corymbose; corolla green, yellow, purplish to reddish brown, the tube neither dorsally cleft nor fenestrate, expanded basally and near throat but with a somewhat constricted waist, the lobes falcate, all bent away from the dorsal side of the flower with the two dorsal lobes the longest and the ventral lobe usually the shortest; filament basally distinct, either basally adherent or free from the corolla, the anther tube with an enlarged, oblique, terminal orifice, the anthers either all terminally naked or some or all of the anthers with a few pilose trichomes or the two lowermost with a dense tangle of villous trichomes but never with a terminal tuft of stiff bristles or a cartilaginous triangular scale; style jointed just above ovary and deciduous with the corolla and stamens, the ovary apically truncate or nearly so. Fruits berrylike, occasionally much inflated at maturity; seeds numerous, oblong or ovoid to fusiform, 0.7-1.3 mm long, much longer than wide, minutely foveate-reticulate and the reticulations elongate parallel to the long axis of the seed.