Cornus L.

Dogwoods (en), Cornouiller (fr)

Genus

Angiosperms > Cornales > Cornaceae

Characteristics

Evergreen or (some cultivated species in Australia, e.g. C. florida) deciduous, shrubs or trees, rarely herbs (e.g. C. canadensis), hairy (in Australia) or glabrous; hairs often medifixed. Leaves opposite (in naturalised C. capitata) or alternate (e.g. C. alternifolia), rarely whorled (e.g. C. canadensis), simple, entire, usually with pinnate venation, mostly distinctly petiolate. Inflorescences terminal involucrate heads (in Australia) or (some cultivated species) branched cymes; bracts large and showy (e.g. cultivated and naturalised C. capitata in Australia), or small or absent (some cultivated species). Flowers bisexual, small, greenish, white or yellow. Sepals 4, minute, calyx-tube campanulate, cylindric or urceolate. Petals 4, oblong, valvate. Stamens 4, alternating with petals; anthers oblong, purple. Ovary usually 2-celled, ovule 1 per cell, style filiform or cylindric, stigma capitate. Fruit a drupe, ovoid, 2-celled, 2-seeded (1 seed in each chamber), in compound heads (in naturalised C. capitata in Australia) or simple; endocarp bony. Seeds oblong, compressed, albumen fleshy.
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Shrubs, trees, or herblike shrubs, precocious, coetaneous, or serotinous. Young shoots pubescent, rarely glabrous; trichomes curly or straight, raised or appressed. Stem sympodial, rarely monopodial. Winter buds terminal or axillary, mixed or separate, covered or exposed. Petiole slightly furrowed adaxially; leaf blade narrowly elliptic, elliptic, oblong, or ovate, glabrous to densely pubescent, lateral veins actinodromous, often raised abaxially. Inflorescence formed in previous or current year; bracts covering inflorescence or not. Sepals 4, fused; teeth absent, minute, or variously triangular. Petals 4, free, spreading, oblong to orbicular, valvate. Filaments filiform or awn-shaped, longer than style, longer or shorter than petals; anthers whitish or yellow, rarely blue, red, or purplish, ellipsoid to narrowly ellipsoid or oblong, 2-loculed. Ovary obovoid, crowned by a disk. Fruit globose, ovoid, oblong, or ellipsoid, crowned by persistent calyx, disk, and style; stones globose, ovoid, ellipsoid, oblong, sometimes asymmetric, surface smooth or ribbed, apex rarely pitted.
Fls mostly perfect, 4-merous; sep usually minute; pet small, valvate; style well developed; stigma capitate; fr a drupe, the stone bilocular (except in C. canadensis) but often only 1-seeded; fls in cymose or cymose-paniculate infls or, by reduction of the branches, in umbelliform or head-like clusters. (Chamaepericlymenum, Svida, Cynoxylon)50+, widespread but mainly N. Temp.Named hybrids include C. ×acadiensis Fernald (C. alternifolia × sericea), C. ×arnoldiana Rehder (C. amomum var. schuetzeana × racemosa), and C. ×slavinii Rehder (C. rugosa × sericea). Others include C. amomum × stricta, C. drummondii × stricta, C. amomum var. schuetzeana × drummondii, C. drummondii × racemosa, and C. racemosa × stricta.
Herbs, shrubs, or trees, clonal from rhizomes, rooting from decumbent branches, or aclonal; hairs 1-celled, arms either short and ornamented with micropapillae and calcium carbonate crystals, or long, erect, curling, and twisted. Leaves: blade lanceolate to broadly ovate; abaxial surface often papillate. Inflorescences: bracts adnate to inflorescence branches, distal portion either minute and caducous or expanding into showy, nonchlorophyllous involucres. Pedicels present or absent. Flowers: hypanthium turbinate or urceolate; petals spreading or recurved, usually cream, rarely purple; stamens exserted; anthers dorsifixed, versatile. Drupes globose, subglobose, or ellipsoid, slightly fleshy. x = 11.
Trees and shrubs, rarely rhizomatous subshrubs. Leaves opposite, rarely alter-nate, estipulate, simple, entire, usually petiolate. Inflorescence an ebracteate corymbiform or umbelliform cyme or (in our species) densely capituliform and subtended by usually 4 involucrate bracts. Flowers small, hermaphrodite, epig-ynous. Sepals 4, small, slightly united at the base. Petals 4, valvate. Stamens 4, the anthers 4-celled, dorsifixed, versatile. Ovary inferior, 2-celled, each cell with a single pendulous ovule; style 1, arising from a fleshy epigynous disc, the stigma minute, capitate. Fruit a drupe, usually containing a single seed.
Life form perennial
Growth form
Growth support -
Foliage retention evergreen
Sexuality hermaphrodite
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Root system rhizome
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Nitrogen fixer -
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Environment

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Hardiness (USDA) 4-11

Usage

Many species and their cultivars are grown as ornamentals, particularly for the white or colourful floral bracts, autumn-coloured leaves, or the bright red or yellow winter-coloured stems of some species. In Australia they are mainly grown in temperate regions, and include e.g. Cornus alba (Red-barked Dogwood, Tatarian Dogwood), C. alternifolia (Pagoda Dogwood), C. canadensis (Creeping Dogwood), C. capitata (Himalayan Strawberry-tree), C. controversa (Table Dogwood), C. florida (Flowering Dogwood, Common White Dogwood), C. kousa (Japanese Flowering Dogwood, Kousa), C. mas (Cornelian Cherry), C. nuttallii (Pacific Dogwood) and C. pumila (Dwarf Dogwood) (Pryor & Banks 1991, Rodd 1996, Ellison 1999, Spencer 2002, Brummitt 2007).                                          Cornus florida with its showy pink and white bracts, cultivated in a garden in Turner, A.C.T. Photo: P.G. Kodela, 12 October 2019. The fruits of Cornus mas are cultivated in Asia for medicinal uses (Xiang & Boufford 2005, Hosseinpour-Jaghdani et al. 2017), and the tannins from C. capitata have been used medicinally (Fern 2019). The wood of some Cornus species is used for a variety of purposes (e.g. timber of C. nuttallii), and some species have edible fruits (e.g. C. mas) used for jams and alcoholic beverages (Spencer 2002: 436; Brummitt 2007: 112).
Uses medicinal ornamental timber wood
Edible -
Therapeutic use -
Human toxicity -
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Cultivation

Mode -
Germination duration (days) 90 - 730
Germination temperacture (C°) 10 - 15
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