Epipactis Zinn

Helleborine (en), Épipactis (fr)

Genus

Angiosperms > Asparagales > Orchidaceae

Characteristics

Herbs, terrestrial, autotrophic or rarely holomycotrophic, medium-sized. Rhizome creeping, short or elongate, with many elongate, fleshy roots. Stem erect, leafy, with 2 or 3 scalelike sheaths toward base, glabrous or pubescent. Leaves 3 to many, cauline, spirally arranged, broadly elliptic to linear-lanceolate, plicate, scabrous or smooth. Inflorescence terminal, racemose, often secund, few to many flowered; floral bracts usually foliaceous. Flowers spreading or nodding, resupinate or very rarely not resupinate, medium-sized. Sepals free, spreading or connivent, glabrous, puberulent, or pubescent. Petals similar to sepals but shorter; lip fleshy, not spurred, constricted near middle to form a distinct epichile and hypochile, and sometimes with a mesochile in between; hypochile cymbiform or saccate, sometimes containing lamellae; mesochile (when present) short or elongate, narrow; epichile spreading, usually hinged at its base, thickened or not, variable in shape, sometimes 3-lobed, sometimes with wartlike calli or keels. Column short, without a column foot; anther sessile; pollinia 4, granular-farinaceous, without caudicles; viscidium absent in autogamous species; rostellum usually large, very rarely absent; stigma rounded to subquadrate. Capsule pendulous or ascending, obovoid to ellipsoid.
More
Terrestrial, occasionally saprophytic herbs with horizontal or vertical, very short rhizomes, numerous fleshy roots and simple, erect, leafy stems. Leaves ovate or lanceolate, plicate, occasionally very small. Flowers in ± secund racemes, rather inconspicuous, spreading or pendulous, shortly pedicelled. Tepals spreading or remaining closed, dull reddish or greenish. Sepals free, subequal. Petals scarcely smaller than sepals. Lip usually in 2 parts articulated by a narrow joint or fold (mesochile); the basal part (hypochile) forms a nectar-containing cup often with a pair of basal bosses; the apical part (epichile) forms a ± cordate or triangular downwardly directed terminal lobe; spur absent. Column short, flat or concave in front with a shallow cup at apex; anther free, hinged at the back of the summit of the column, behind the stigma and rostellum, ovate, slightly convex, 2-celled; pollinia 2, tapering towards their apices near which they are attached to the rostellum, each ± divided longitudinally into halves; caudicles absent; pollen grains forming friable masses loosely bound by fine threads; stigma prominent, broad; rostellum placed centrally above the stigma, large, globular, persistent, evanescent or absent. Capsule oblong, spreading or pendulous.
Herbs perennial, terrestrial, glabrous or puberulent to glabrate. Roots from short rhizome, fibrous. Stems leafy. Leaves several, alternate, sheathing, plicate, grading into floral bracts. Inflorescences terminal racemes; floral bracts leaflike, often exceeding flowers. Flowers few to many, resupinate, pedicellate; perianth free, spreading or loosely connivent, subequal; sepals ovate to ovate-lanceolate, concave; petals often shorter than sepals; lip sessile at base of column, fleshy, divided into 2 parts by central constriction, proximal part (hypochile) deeply concave to saccate, distal part (epichile) triangular to oblanceolate, with pair of calli near base; column curved over lip, broadened distally, short; anther terminal, sessile; pollinia 4, mealy. Fruits capsules, pendent to spreading, obovoid to ellipsoid.
Sep and lateral pet similar; lip strongly saccate in the basal half, the terminal lobe broadly ovate, not hinged to the basal part, crested at its base with 2 evident swellings; column very short and broad, the large sessile anther hinged to its back below the middle and somewhat projecting; pollinia 4; rhizomatous or stoloniferous plants with numerous alternate lvs and a terminal raceme. 25, N. Hemis.
Column short, flat or concave in front, with a shallow cup at the apex; anther free, 2-celled, hinged at the back of the column apex, behind the stigma and rostellum; pollinia 2, attached near the rostellum apex, each longitudinally divided, caudicles absent; pollen grains in friable masses loosely bound by fine threads.
Lip with no spur, but with the hypochile forming a nectar-containing cup; hypochile articulated to the epichile by a narrow joint.
Terrestrial herb, occasionally saprophytic, with very short rhizomes, numerous fleshy roots and simple erect leafy stems.
Inflorescence racemose; flowers usually dull reddish or greenish, pedicelled.
Stigma broad, with the large, globular rostellum placed centrally above it.
Leaves ovate or lanceolate, plicate.
Sepals and petals free.
Life form perennial
Growth form herb
Growth support -
Foliage retention -
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) -
Root system creeping-root fibrous-root rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 4-11

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Epipactis unspecified picture
Epipactis unspecified picture

Distribution

Epipactis world distribution map, present in Afghanistan, Albania, Austria, Burkina Faso, Bulgaria, Belarus, Switzerland, Chile, China, Colombia, Cyprus, Czech Republic, Germany, Denmark, Algeria, Spain, Ethiopia, Finland, France, Micronesia (Federated States of), Greece, Hungary, Indonesia, Ireland, Iran (Islamic Republic of), Iraq, Italy, Japan, Kazakhstan, Kenya, Kyrgyzstan, Lao People's Democratic Republic, Morocco, Myanmar, Montenegro, Mozambique, Malaysia, Nicaragua, Netherlands, Norway, Nepal, Oman, Pakistan, Palau, Poland, Korea (Democratic People's Republic of), Portugal, Romania, Russian Federation, Rwanda, Saudi Arabia, Singapore, Somalia, Sweden, Turks and Caicos Islands, Thailand, Turkmenistan, Taiwan, Province of China, Tanzania, United Republic of, Uganda, Ukraine, United States of America, Uzbekistan, Viet Nam, and Yemen

Identifiers

LSID urn:lsid:ipni.org:names:329384-2
WFO ID wfo-4000013619
COL ID 8VXDW
BDTFX ID 86382
INPN ID 192235
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Amesia Epipactis

Lower taxons

Epipactis nordeniorum Epipactis autumnalis Epipactis exilis Epipactis rivularis Epipactis atromarginata Epipactis atrorubens Epipactis x conquensis Epipactis x gerbaudiorum Epipactis gigantea Epipactis greuteri Epipactis guegelii Epipactis liestalensis Epipactis olympica Epipactis phyllanthes Epipactis placentina Epipactis x reinekei Epipactis royleana Epipactis x schmalhausenii Epipactis x schulzei Epipactis thunbergii Epipactis tremolsii Epipactis veratrifolia Epipactis x vermionensis Epipactis xanthophaea Epipactis x robatschii Epipactis x hanseniorum Epipactis x bruxellensis Epipactis bugacensis Epipactis x cardonneae Epipactis fageticola Epipactis flaminia Epipactis graberi Epipactis x heterogama Epipactis kleinii Epipactis lusitanica Epipactis meridionalis Epipactis albensis Epipactis barlae Epipactis x barreana Epipactis x breinerorum Epipactis condensata Epipactis x gevaudanii Epipactis ohwii Epipactis papillosa Epipactis pontica Epipactis x pupplingensis Epipactis x soguksuensis Epipactis x stephensonii Epipactis tallosii Epipactis troodi Epipactis turcica Epipactis turcomanica Epipactis persica Epipactis babianifolia Epipactis persica Epipactis stellifera Epipactis cupaniana c.brullo, d'emerico Epipactis krym-montana Epipactis sanguinea Epipactis etrusca Epipactis calabrica Epipactis hyblaea Epipactis pinovica Epipactis fascicularis Epipactis x mohilewiensis Epipactis dunensis Epipactis humilior Epipactis collaris Epipactis africana Epipactis flava Epipactis mairei Epipactis ulugurica Epipactis rechingeri Epipactis alatus Epipactis persica Epipactis ioessa Epipactis x nicolosii Epipactis leptochila Epipactis microphylla Epipactis purpurata Epipactis muelleri Epipactis palustris Epipactis helleborine