Holcus L.

Velvetgrass (en), Houlque (fr)

Genus

Angiosperms > Poales > Poaceae

Characteristics

Perennials or rarely annuals, rhizomatous to stoloniferous or caespitose, bisexual. Leaves: ligule an unfringed or fringed membrane; blade rolled in bud. Inflorescences paniculate, loose or ±contracted. Spikelets pedicellate, laterally compressed, falling with glumes, 2-flowered, with lower floret bisexual and upper floret male-only or sterile or very rarely bisexual; rachilla usually with a distinctly elongated internode above glumes and between florets, with lower floret elevated on a markedly curved internode, prolonged above upper floret, extension sometimes with an incomplete floret. Glumes 2, ±equal or upper glume sometimes somewhat longer, ±equalling to exceeding florets and enclosing them, awned or only upper glume awned or awnless, keeled; lower glume 1-nerved; upper glume 3-nerved. Callus very short. Basal lemma awnless or with 1 dorsal geniculate awn (not in Australia), decidedly firmer than glumes, leathery or cartilaginous, keeled or dorsally rounded, indistinctly 3–5-nerved, shiny. Palea tightly clasped by lemma, textured like lemma, 2-keeled. Lodicules membranous. Stamens 3. Caryopsis laterally compressed; hilum short or rarely long-linear; embryo small. Upper floret awn 1, dorsal, conspicuous, short, arising from above middle of lemma, curved or hooked or geniculate.
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Annuals or perennials. Leaf blades flat. Inflorescence a moderately to densely contracted panicle. Spikelets laterally compressed, disarticulating below glumes, florets 2, lower floret bisexual, upper floret staminate; rachilla usually curved and hooked below bisexual floret, often shortly extended above staminate floret; glumes subequal, papery, enclosing florets, strongly keeled, lower glume 1-veined, upper glume 3-veined; lemmas cartilaginous, shiny, rounded on back, veins indistinct, apex obtuse or 2-toothed, upper lemma or both awned; awn arising from upper 1/3 of lemma back, geniculate, hooked or straight; palea slightly shorter than lemma, membranous.
Spikelets mostly 2-fld, disarticulating below the glumes; glumes thin, keeled, much exceeding and concealing the lemmas, the first 1-veined, the second broader, 3-veined; lemmas rounded on the back, obscurely veined, the lower awnless (in most spp., incl. ours), enclosing a perfect fl, the upper awned, with a staminate fl; awn arising below the lemma-tip; perennials (ours) with flat, soft, relatively wide lvs and dense, contracted, ovoid or subcylindric panicles of small spikelets. 8, Old World.
Life form perennial
Growth form herb
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Sexuality hermaphrodite
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Hardiness (USDA) 4-10

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