Oryza L.

Rice (en), Riz (fr)

Genus

Angiosperms > Poales > Poaceae

Characteristics

Annuals or perennials, caespitose, rhizomatous or stoloniferous, bisexual. Culms herbaceous. Leaves usually auriculate; ligule an unfringed membrane, often lacerated; blade without cross-nerves. Inflorescences paniculate, open or sometimes contracted, with axes usually wavy; primary branches often simple and raceme-like. Spikelets pedicellate, disarticulating above glumes or rarely persistent, strongly laterally compressed, with 2 basal florets reduced to sterile lemmas and 1 terminal bisexual floret; pedicel apex cupuliform. Glumes rudimentary, minute, scale-like or reduced to a rim represented by a small 2-lobed cupule. Sterile lemmas equal (or slightly unequal), c. ¼–½ or less length of spikelet. Bisexual floret: lemma firmer than sterile lemmas, awnless, mucronate to cuspidate or with an antrorsely scabrous, straight to somewhat sinuous terminal awn, strongly keeled, predominantly (3–) 5-nerved, incurled, clasping lateral nerves of palea, closely papillose or colliculate, scabrous to hispid especially (or only) on nerves, the stiff hyaline hairs or spines prominent and often longer towards apex (glabrous in some species outside Australia); palea textured like lemma, 3-nerved; stamens usually 6. Caryopsis laterally compressed or terete; hilum long-linear.
More
Annual or perennial, tufted or shortly rhizomatous. Culms erect or ascending. Leaf blades mainly cauline, broadly linear, flat; ligule membranous, sometimes long. Inflorescence a panicle, usually many-spiculate, often nodding, lower branches usually whorled, unbranched or sparsely branched, pedicels short. Spikelets with 3 florets, 2 lower florets reduced, sterile, upper floret fertile, strongly laterally compressed, disarticulating below sterile lemmas, persistent in cultivated species; glumes vestigial, remaining after disarticulation as a shallow lobed frill at pedicel apex; sterile florets reduced to 2 narrow lemmas at base of fertile floret; fertile lemma boat-shaped, keeled, leathery, closely papillose, sometimes spinulose, infrequently smooth, prominently 5-veined, apex awnless to long awned; awn straight; palea resembling lemma but narrower, 3-veined, apex beaked. Stamens 6. Caryopsis variable in shape, embryo 1/4 length of caryopsis, hilum linear, as long as caryopsis. x = 12.
Erect annuals or perennials with flat leaf-blades and membranous ligules. Panicle open or rather dense; branches numerous, racemose with shortly pedicelled spikelets. Spikelets 3-flowered, strongly laterally compressed, falling entire from the pedicel; glumes absent or represented by obscure lips at the pedicel-tip; both lower florets represented only by narrow lemmas, these usually subulate or linear and much shorter than the spikelet. Upper floret hermaphrodite; lemma coriaceous, strongly keeled, prominently 5-nerved, with or without an awn; palea similar but narrower, keeled, 3-nerved; stamens 6; stigmas 2.
Spikelets 1-fld, articulated below the glumes, laterally compressed; glumes subequal, triangular-subulate, 1-veined, much shorter than the lemma; lemma narrowly oblong at anthesis, boat-shaped at maturity, acute, awned or awnless, 5-nerved, the lateral nerves near the involute margins; palea about equaling the lemma; plants of wet soil, with wide flat lvs and large open panicle. 25, trop.
Fertile lemmas 5-nerved (one pair of nerves very close to the margins), cymbiform, strongly keeled, coriaceous or rarely chartaceous, often tessellate or rugulose, usually awned, rarely awnless (in cultivated rice), usually with 2 short lateral protrusions at the apex; awn usually much longer than the spikelet, sometimes coloured, stiff or flexuous, often with a coloured basal callus.
Spikelets 1-flowered, disarticulating below the glumes, laterally compressed; glumes narrow, much shorter than the lemma (in our species); lemma indurate, rigid, keeled, 3-nerved, awned or sometimes awnless, sparsely to rather densely hispid; palea similar to the lemma but narrower and with no midnerve on the back, the two nerves close to the margins.
Palea slightly shorter than the lemma, of the same consistency and indument but with the margins hyaline, 3-(rarely 5-) nerved, awnless, obtuse or acute to acuminate, usually tightly clasped by the inflexed margins of the lemma.
Caryopsis (usually firmly enclosed by lemma and palea thus forming the “husk”), laterally compressed, oblong to oblong-elliptic in outline, whitish to brown or reddish; embryo very small; hilum large, linear.
Spikelets shortly pedicelled, 1-flowered, solitary, laterally strongly compressed, deciduous or (in cultivated rice) persistent, hermaphrodite; rhachilla articulating above the rudimentary glumes.
Sterile lemmas 2, subequal in shape and size, usually very small (nearly as long as the spikelet only in one sp. from S. America), faintly 1-nerved or nerveless.
Glumes 2, reduced to a narrow entire or 2-lobed rim, forming a cup-shaped cavity at the apex of the pedicel, whitish, green or reddish in colour.
Culms few-to many-noded, erect or ascending, sometimes floating, often spongy, usually hollow between the nodes.
Inflorescence an open or contracted panicle, many-flowered, erect or drooping.
Annual or perennial grasses, growing in a variety of moist habitats.
Ligule membranous, often lacerate, glabrous or pilose.
Lodicules 2, small, obovate or lanceolate-elliptic.
Stigmas pilose, large, laterally exserted.
Stamens 6, with the filaments free.
Ovary glabrous.
Life form
Growth form herb
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Foliage retention deciduous
Sexuality hermaphrodite
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Hardiness (USDA) 8-12

Usage

Rice, especially Oryza sativa (commonly known as Asian Rice) but also another domesticated species, O. glaberrima (African Rice), is an economically important cereal grain crop and staple food in many parts of the world. It is consumed by a large part of the world's population, especially in Asia, and is one of the world's most highly produced agricultural commodities. Australian native species, e.g. Oryza australiensis, have been used in genetic and hybrid rice technology studies. As discussed by Henry et al. (2010) the Australian indigenous species have potential for domestication in some cases and have many traits of potential value for the improvement of domesticated rice. Stress tolerance (biotic and abiotic) and grain quality characteristics in the Australian populations may be useful (Henry et al. 2010). Wurm et al. (2012) investigated the milling and processing, mineral nutritional and putative cooking qualities of rice grain samples from wild populations of the Australian native species O. rufipogon and O. meridionalis; to find their suitability for bush-tucker, novelty or other wild rice enterprises. See also Henry (2019), Abdelghany et al. (2022) and references therein. "Australian wild Oryza species have potential as high-value, low-volume, culturally identified, and nutritious food, especially in gourmet food, tourism, restaurants, and value-added products" (Abdelghany et al. 2022). In 2019, trials were planned to cultivate wild rice near Darwin to establish a potential native rice industry in Australia (ABC News report: https://www.abc.net.au/news/rural/2019-04-02/native-rice-trials-set-for-the-northern-territory/10959786; 2 April 2019). An earlier ABC Science article by A. Thomas (15 October 2010):   http://www.abc.net.au/science/articles/2010/10/15/3038568.htm In addition to rice being a major food source (as whole grain, flour, rice milk and in processed foods) it is also used in medicinal treatments (the grain and rhizome), the grain is fermented to produce rice vinegar and wine (e.g. Japanese sake), there are uses for rice starch (e.g. in cosmetics and textiles), oil from the seed (e.g. for cooking and in soap and plastic materials), rice straw and husks (e.g. thatched roofing, stock feed, ropes, weaving (e.g. mats, hats, bags), packaging, paper production and fuel, especially in Asia and Africa), and rice grains are used in re-usable heat packs. Rice plays an important role in many cultures and features in history and design.
Uses medicinal oil
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Cultivation

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