Schoenus L.

Bogrush (en), Choin (fr)

Genus

Angiosperms > Poales > Cyperaceae

Characteristics

Perennial tufted herbs, 0.1–0.55 m high; plants clonal, usually forming dense tussocks 0.05–0.2 m across; rhizome thin, woody, short, branched, 0.8–3.4 mm diam., pseudobulbous; old leaf sheaths covering the rhizome, usually breaking apart into fine fibres with age, straw-coloured to reddish brown; roots sand-binding or not. Culms slender, rigid, erect, 11–53 cm tall, 0.6–2.5 mm wide, not noded, terete to compressed, usually sulcate when dried, smooth, glabrous, green to yellow-green, base not enlarged. Leaves all basal, numerous; phyllotaxy loosely spirotristichous; sheath 20–70 mm long, 0.7–2.7 mm wide, broader than the leaf lamina, glabrous or scabrid, straw-coloured to reddish brown, dull, upper margin membranous, oblique, ligule absent; cauline sheath tight around culm; pseudopetiole obscure or absent; leaves with a well-developed lamina, not exceeding the culms, lamina dorsiventral, 4–21 cm long, 0.3–0.9 mm wide; bases white or brown-membranous, sometimes dividing; lamina ± linear, flat to channelled above, ± flexuous, old leaf tips usually curling, finely multi-striate, glabrous except for margins, green to brown, concolorous, keeled along mid-vein or not, margins scaberulous, not recurved, apex long-attenuate, setaceous; lowest leaves reduced to a sheath, lamina reduced to a mucro, pale to reddish brown. Inflorescence very contracted, panicle-like or subcapitate and appearing head-like, straight to somewhat flexuous, with 2–6 nodes, 0.5–1.7 cm long, 6–16 mm wide, not interrupted; bracts leaf-like, greatly exceeding the spikelet clusters in length, sheaths in the lower part of the inflorescence open, not or sometimes enveloping the higher bracts, gradually reducing along the inflorescence and often mostly hidden among the spikelets, basal bract spreading to suberect, similar to the leaves, much longer than the inflorescence, to 65 mm long; branches erect to spreading, with 2–5 spikelets in a terminal cluster, spikelet(s) on short, terete, glabrous peduncles 0.2–1.4 mm long, arising in each bract axil; basal branchlet 4.5–6.6. mm long (including spikelets); spikelet prophyll present, sheath 0.9–1.5 mm long, brown, partially enclosing the spikelet, with a slender lamina 0.2–1.6 mm long, brown. Spikelets pale to dark brown or chestnut, narrow-ovate, 4.7–8 mm long, 1.2–3.2 mm wide, terete, with 5–9 glumes, only topmost 1 or 3 fertile, the lower 2 flowers when present male fertile, the upper flower bisexual, fertile, no reduced glume above the florets; glumes spirodistichous, basal glume 2.6–4.2 mm long, fertile glumes 3.8–6.2 mm long, 0.9–1.6 mm wide, membranous, pale to dark brown or chestnut, lamina puberulent or glabrous, keel and margins scaberulous, ciliate or denticulate, acute or attenuate, often sparsely scabrous on the keel, usually ciliate on the margins, lamina glabrous, ovate-lanceolate, acute to acuminate or obtuse, with a mucro 0.3–1.2 mm long, keel distinct; rachilla compact and not sinuous in fruiting spikelets. Perianth segments 6, whitish, minute, ± linear, flat to terete, margins with dense to scattered, white, short, antrorse, ciliate or ciliate-plumose hairs, persistent on the nutlet. Stamens 3; anther connective 2.0–6.2 mm long, linear to subulate; anthers yellow, 1.8–3.1 mm long, twisted when dry, with a glabrous apical appendage 0.5–0.8 mm long. Style trifid, base 1.4–3.5 mm long, glabrous, slender throughout, branches 1.3–2.9 mm long, hispidulous. Nutlet stipitate, stipe 0.3–0.5 mm long, somewhat constricted, obovoid to obpyriform, obscurely trigonous, dull, brown to dark brown, reddish brown or grey, 1.4–2.3 mm long including the stipe and style base, 1.0–1.4 mm diam., faces irregularly reticulate to rugulose or tuberculate at 40× magnification, shortly hispidulous to scabrous or tuberculate at the apex, otherwise glabrous; epidermal cells usually square–hexagonal, inconspicuous, with three fine white ribs, style base not enlarged, thin, sometimes part of the base persistent, cylindrical; embryo Schoenus-type. Photosynthetic pathway inferred from anatomy to be C3. 1C = 0.26 pg for C. turbinata (K.L. Wilson 10781; Bureš et al. 2013).
More
Perennials with woody, abbreviated or shortly creeping rhizome, more rarely annuals. Stems erect, or ascending and rooting at the nodes, terete or obtusely trigonous, striate or sulcate, usually smooth. Leaves either all basal, or basal and cauline, linear, often setaceous, canaliculate, sometimes reduced to mucronate sheaths; ligule absent; basal sheaths open, dark-coloured; sheaths of the cauline leaves closed, tubular, often bearded at the mouth. Inflorescence terminal, racemose or paniculate, consisting of some distant fascicles of branches subtended by a leafy bract, or contracted and head-like. Spikelets solitary or clustered, (in the Malesian spp.) compressed, lanceolate to oblong-lanceolate, sometimes falcate, usually few-flowered. Rachilla straight and with very short internodes in the lower part (between the empty glumes), the upper internodes (between the fertile glumes) elongated and prominently zigzag. Glumes distichous, acropetally caducous, 1-nerved, keeled, frequently dark-coloured, the lower ones empty, the upper (flower-bearing) ones usually decurrent on the rachilla ('rachilla winged'), the terminal one often strongly reduced and empty. Flowers in the hollows of the zigzag rachilla, usually bisexual, but the uppermost often more or less reduced (male or functionally male). Perigone consisting of up to 6 filiform or linear-lanceolate, ciliate to plumose or antrorsely scabrous bristles, or absent. Stamens (1-)3(-6), (in Mal. (2-)3); anthers linear, with more or less produced glabrous connective. Style slender, continuous with the ovary, not or hardly dilated at the base, brown, caducous (often the very base remaining on the nut as an apiculus not separated from the nut proper by a constriction). Stigmas 3 (very rarely 2). Nut sessile or shortly stipitate, ovoid, obovoid, or ellipsoid, trigonous, often 3-ribbed, very rarely biconvex, glabrous, or hispidulous in the upper part, smooth, rugulose, or scrobiculate; epidermal cells isodiametric to oblong.
Infl. paniculate, or capitate, or a solitary spikelet. Spikelets 1–4-fld, sessile or stalked, fls hermaphrodite, or the uppermost functionally male; rhachilla flexuous or zig-zag between the fls; glumes distichous, keeled, us. nerveless except for a central midrib, some lower glumes and some upper ones often empty. Hypog. bristles 6–1, plumose or ciliate, us. scabrid; or 0. Stamens 3, occ. 2. Style-branches 3, occ. 2. Nut ± trigonous, angles ± thickened. Perennial herbs, with or without creeping rhizomes, or rarely annual. Culms erect, curved, or drooping and rooting below, terete or compressed, branched or unbranched. Lvs basal or cauline, narrow-linear, setaceous and rigid, or flaccid, margins of lamina us. slightly toothed; or all, or the basal lvs reduced to sheathing, us. mucronate bracts. About 100 spp. of temperate regions. Of the 8 N.Z. spp., 2 are endemic; the other 6 occur in Australia as well and 3 of these are recorded from elsewhere in the Pacific.
Life form perennial
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
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Mature height (meter) 0.1 - 0.55
Root system rhizome
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Nitrogen fixer -
Photosynthetic pathway -

Environment

In Malesia the members of sect. Calostachyi, Paniculati, and Nudicaules are mainly inhabitants of savannahs and savannah forests in the periodically dry regions; the species of sect. Helothrix occur only at high altitudes.
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Hardiness (USDA) 6-12

Usage

Uses -
Edible -
Therapeutic use -
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Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
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Images

Schoenus unspecified picture
Schoenus unspecified picture

Distribution

Schoenus world distribution map, present in Albania, Australia, Austria, Bulgaria, Bahamas, Belarus, Belize, Switzerland, Cuba, Cyprus, Czech Republic, Germany, Dominica, Denmark, Algeria, Eritrea, Spain, Ethiopia, Finland, Fiji, France, Micronesia (Federated States of), Greece, Honduras, Hungary, Ireland, Iran (Islamic Republic of), Iraq, Italy, Jamaica, Japan, Kyrgyzstan, Libya, Lesotho, Morocco, Moldova (Republic of), Nicaragua, Netherlands, Norway, New Zealand, Pakistan, Philippines, Palau, Papua New Guinea, Poland, Portugal, Romania, Russian Federation, Saudi Arabia, Singapore, Svalbard and Jan Mayen, Solomon Islands, Somalia, Sweden, Turks and Caicos Islands, Chad, Thailand, Turkmenistan, Tunisia, Taiwan, Province of China, Ukraine, Uruguay, United States of America, Uzbekistan, Viet Nam, Wallis and Futuna, Yemen, Zambia, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:329458-2
WFO ID wfo-4000034547
COL ID 8VSG9
BDTFX ID 87177
INPN ID 197469
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Schoenus Chaetospora

Lower taxons

Schoenus asperocarpus Schoenus badius Schoenus biglumis Schoenus centralis Schoenus clandestinus Schoenus compar Schoenus curvulus Schoenus deformis Schoenus dregeanus Schoenus efoliatus Schoenus elegans Schoenus ericetorum Schoenus falcatus Schoenus ferrugineus Schoenus grandiflorus Schoenus griffinianus Schoenus laevigatus Schoenus laevinux Schoenus lanatus Schoenus latitans Schoenus lepidosperma Schoenus odontocarpus Schoenus pauciflorus Schoenus plumosus Schoenus smitinandii Schoenus subaphyllus Schoenus subflavus Schoenus sublateralis Schoenus sublaxus Schoenus tenuissimus Schoenus trachycarpus Schoenus unispiculatus Schoenus yarrabensis Schoenus absconditus Schoenus antarcticus Schoenus apogon Schoenus caespititius Schoenus calostachyus Schoenus capillifolius Schoenus carsei Schoenus cruentus Schoenus curvifolius Schoenus cuspidatus Schoenus delicatulus Schoenus evansianus Schoenus falcatus Schoenus fluitans Schoenus latelaminatus Schoenus longibracteatus Schoenus lymansmithii Schoenus natans Schoenus neocaledonicus Schoenus obtusifolius Schoenus pennisetis Schoenus pleiostemoneus Schoenus pygmaeus Schoenus quadrangularis Schoenus racemosus Schoenus scabripes Schoenus x scheuchzeri Schoenus sculptus Schoenus subbarbatus Schoenus subfascicularis Schoenus tesquorum Schoenus variicellae Schoenus villosus Schoenus achaetus Schoenus acuminatus Schoenus andinus Schoenus andrewsii Schoenus armeria Schoenus benthamii Schoenus bifidus Schoenus brevifolius Schoenus brevisetis Schoenus discifer Schoenus globifer Schoenus grammatophyllus Schoenus hexandrus Schoenus humilis Schoenus imberbis Schoenus indutus Schoenus insolitus Schoenus kennyi Schoenus loliaceus Schoenus maschalinus Schoenus melanostachys Schoenus minutulus Schoenus moorei Schoenus nanus Schoenus multiglumis Schoenus nitens Schoenus paludosus Schoenus pedicellatus Schoenus punctatus Schoenus rhynchosporoides Schoenus rigens Schoenus rodwayanus Schoenus setiformis Schoenus sinensis Schoenus sparteus Schoenus subbulbosus Schoenus submicrostachyus Schoenus tendo Schoenus tenellus Schoenus turbinatus Schoenus calostachyus Schoenus rivularis Schoenus thedae Schoenus tenuellus Schoenus adnatus Schoenus albovaginatus Schoenus arenicola Schoenus aureus Schoenus auritus Schoenus bolusii Schoenus bracteosus Schoenus calceolus Schoenus comparoides Schoenus complanatus Schoenus crassiculmis Schoenus crassus Schoenus exilis Schoenus filiculmis Schoenus galpinii Schoenus gracillimus Schoenus graminifolius Schoenus lucidus Schoenus megacarpus Schoenus neovillosus Schoenus prophyllus Schoenus pseudoloreus Schoenus purpurascens Schoenus quartziticus Schoenus riparius Schoenus schonlandii Schoenus submarginalis Schoenus triticoides Schoenus autumnalis Schoenus variabilis Schoenus compactus Schoenus ornithopodioides Schoenus vaginatus Schoenus calyptratus Schoenus cygneus Schoenus breviculmis Schoenus calcatus Schoenus nudifructus Schoenus pictus Schoenus sesquispicula Schoenus nigricans