Shrubs or trees, dioecious or monoecious. Young persistent branchlets usually distinguishable from the deciduous branchlets. Branchlet articles with furrows deep, concealing the stomates. Leaves in whorls of 4–14. Male flowers in simple short to long spikes, on branchlets which in the flowering region are usually distinctly different from the vegetative branchlets. Female inflorescences on short lateral branches or ± sessile. Infructescences ('cones') pedunculate or sessile; bracts thin in exposed portion; bracteoles considerably thickened and often divided so that dorsal portion forms 1 or more distinct protuberances. Samara body brown to black, shining, glabrous or hairy.
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